Histone in nucleosome core are all except
In addition to binding to the nucleosome, the H1 protein binds to the "linker DNA" (approximately 20-80 nucleotides in length) region between nucleosomes, helping stabilize the zig-zagged 30 nm chromatin fiber Ref-Harpers illustrated biochemistry 30/e p280
H2A
H2B
H3
H1
Metabolism of nucleic acids