Cylindropuntia californica
Cylindropuntia californica is a species of cholla cactus known by the common name snake cholla. It is primarily found in Baja California, Mexico and the southernmost part of California in the United States. It is characterized by a short, decumbent habit, yellow-green flowers, elongated stems, and short spines. It is mostly found in coastal sage scrub and coastal chaparral habitats, but two varieties in Baja California can be found in foothills and deserts. In California, variety californica is regarded as a rare and threatened plant, with a California Native Plant Society listing of 1B.1, in part due to its limited number of occurrences and threats from development. It formerly was considered to have a larger range due to the inclusion of Cylindropuntia bernardina within it as the variety parkeri. Description A short, sprawling cactus, this species is typically found growing less than high. There are usually several to many trunks, with the main branches and stems usually sprawling along the ground or becoming erect. The terminal segments of the stems are usually less than long and measure in diameter. The tubercles on the stem measure and are less than high. The spines of this cactus are generally less than long, and are colored a yellow to orange-brown. Most species of cholla have paper-like sheath enclosed over their spines. On this species, the sheath is a translucent white to gold-brown. The flower is yellow to yellow-green, and the filaments are green. The fruit is leathery, and may have no spines or many. The seeds are less than long. Taxonomy Taxonomic history This species was first detailed by English botanist Thomas Nuttall, who arrived in San Diego in May 1836 by ship. He noted the small yellow flowers, numerous clusters of long and short spines, and arid habitat. He sent this description to John Torrey and Asa Gray under the name Cactus californicus. Variety delgadilloana was originally described in 2001 by Jon Rebman and Donald Pinkava as Cylindropuntia delgadilloana. It is named after the botanist Jose Delgadillo. Variety rosarica was originally described by George Lindsay in 1942 as Opuntia rosarica, with the type material collected by Lindsay at San Telmo in August 1941. Varieties The infraspecific taxa of Cylindropuntia californica are variously recognized as subspecies or varieties. The list below follows the treatment in the Jepson eFlora and the 2016 Annotated Checklist of the Vascular Plants of Baja California, Mexico produced by the San Diego Natural History Museum. Cylindropuntia californica (Torr. & A.Gray) F.M.Knuth var. californica —The autonymic subspecies commonly known as the snake cholla. Found sparsely around San Diego, California and northern Baja California. Chromosome number 2n = 22 Cylindropuntia californica (Torr. & A.Gray) F.M.Knuth var. delgadilloana (Rebman & Pinkava) Rebman — Commonly known as the Delgadillo cholla. An erect, robust plant with multiple trunks high. The stem segments are long by in diameter. There are 5 to 8 spines per areole, with 1 to 4 central ones and 3 to 5 radial spines. There are sometimes up to 3 sheathless