Sphincterochila zonata zonata
Sphincterochila zonata zonata is a subspecies of air-breathing land snail, a terrestrial pulmonate gastropod mollusk in the family Sphincterochilidae. This species lives in deserts in Israel and Egypt. Helix boissieri is the type species of the genus Sphincterochila. The type species was subsequently designated by Henry Augustus Pilsbry in 1895. It is named after botanist Pierre Edmond Boissier. Sphincterochila zonata zonata lives in the Negev desert (Israel), and the Sinai desert in the Sinai Peninsula (Egypt). Shell description Shell is cretaceous, white; consists of five convex whorls and a deflected ultimate whorl. The aperture is thickened, projecting internally in two subconcrescent denticles (described by Tryon as "tubercles"). The average diameter of the shell is 25 mm. Anatomy and physiology The average body mass of Sphincterochila zonata zonata is around 4.3 g. Such mass is approximately divided in an even manner between the shell itself and the animal's soft parts. Schmidt-Nielsen et al., in 1971, found that 56% of the animals body mass was contained in its shell. About 80 to 90% of the mass of the soft body parts is composed of water (according to the Yom-Tov 1970). In average, soft body parts contain 81% of water (nearly 1400 mg of water), 11% of proteins, 4% of ash, and little other organic matter. Sphincterochila zonata zonata has no energy reserves and the amount of lipids it contains is a fraction of 1%, which is considered to be extremely low. Ecology Habitat Sphincterochila zonata zonata lives in desert environments. This snail is common in areas with loess-limestone soils, and uncommon in areas that have a flint substrate. Yom-Tov measured the maximum demographic density for Sphincterochila zonata zonata, encountering a value of 0.2-0.3 specimens/m2 in the area of the Negev desert he investigated in 1970. The snail Xerocrassa seetzeni was found to be more abundant there. On the other hand, in the Northern Negev area investigated by Steinberger et al. In 1981 Sphincterochila zonata zonata was the most abundant snail. Sphincterochila zonata zonata along with other snail species and algae are the most significant faunal and floral components of the Negev and Sinai deserts ecosystems. Adaptations for arid conditions This species presents adaptations to arid conditions which significantly improve its desiccation tolerance. Some of those adaptations include a thick shell and a relatively reduced aperture (see also Machin 1967), a thick epiphragm, and slow body surface heat conduction. About 90% of its shell surface reflects the visible portion of the solar spectrum, and much over 90% of the solar spectrum itself. Sphincterochila zonata zonata also produces a new epiphragm after every period of activity (see also Yom-Tov 1971). These snails dig themselves into the soil to depths from 1 to 5 cm while they aestivate during summer in the Negev Desert. In the vicinity of the Dead Sea, they usually either burrow to depths of up to 10 cm, or aestivate hidden under stones. All dormant snails of this species can resist ambient temperatures up to 50 °C, but temperatures of 55