ðə tˈɛɹɪtˌoːɹi ɪz mˈɑːɹkt baɪ bˈoʊθ sˈɛksᵻz, æz ðeɪ bˈoʊθ hæv dɪvˈɛləpt ˈeɪnəl ɡlˈændz fɹʌm wˈɪtʃ ðeɪ ɛkstɹˈuːd ɐ blˈæk sˈʌbstəns ðæt ɪz smˈɪɹd ˌɔn ɹˈɑːks ɔːɹ ɡɹˈæs stˈɔːks ɪn lˈɔŋ stɹˈiːks.
The territory is marked by both sexes, as they both have developed anal glands from which they extrude a black substance that is smeared on rocks or grass stalks in -long streaks.