The American biologist Sewall Wright, who had a background in animal breeding experiments, focused on combinations of interacting genes, and the effects of inbreeding on small, relatively isolated populations that exhibited genetic drift.
Američki biolog Sewall Wright, koji je imao podlogu u eksperimentima križanja životinja, fokusirao se na kombinacije međudjelujućih gena i učinke križanja na malim, relativno izoliranim populacijama koje su pokazivale genetski pomak.