Ludodactylus [SEP] parent taxon
The skull would have been more than 66 cm (26 in) long.
Classification.
Frey "et al." in 2003 classified "Ludodactylus" in the family Ornithocheiridae. In 2007 Frey, had reconsidered the validity of "Ludodactylus", suggesting that it may represent the same animal as, and be a junior synonym of, "Brasileodactylus". However, Andres & Myers (2013), in a large cladistic analysis of pterosaurs, found "Ludodactylus" to be slightly more closely related to
hollow interiors. The axis bears a spiked spine.
Phylogeny.
"Guidraco" was by the describers assigned to the Pteranodontoidea "sensu" Kellner. A phylogenetic analysis found it to be the sister taxon of the Brazilian "Ludodactylus", the two species together forming a clade that was closely related to the Istiodactylidae and the Anhangueridae. The fact that a Chinese form is closely related to a South-American species would indicate a large faunal interchange between continents in this period. This result was later replicated by Wang et