Replication. In cells where centrioles are present as a pair, replication takes place during the whole of the cell cycle. In phase G1 the two centriole cylinders move very slightly apart from one another. During S phase new cylinders of microtubules form near, and at right angles to, the two âmotherâ cylinders.entrioles are absent from the cells of higher plants. When animal cells undergo mitosis they are considered by some to benefit from the presence of centrioles which appear to control spindle fibre formation and which later has an effect on chromosome separation.