sentence: there are four recognized subspecies : cynopterus horsfieldii horsfieldii - java and islands east to sumbawa cynopterus horsfieldii harpax - thailand , peninsular malaysia , sumatra cynopterus horsfieldii persimilis - borneo cynopterus horsfieldii princeps - nias biology and behavior horsfield 's fruit bats eat the fruit of strangler fig , elaeocarpus , and payena , and the flowers of bitter bean .