Vacuolar transporter that regulates zinc homeostasis by mediating zinc transport and storage into the vacuole. ZRC1 senses zinc availability in the cytosol, which might be performed through the histidine repeat motifs, and transports zinc from the cytosol to the vacuole if zinc in cytosol is abundant, conferring resistance to zinc toxicity. Plays a role in resistance to zinc shock resulting from sudden influx of zinc into cytoplasm when ZRT1 and ZRT2 are induced in response to zinc depletion