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In a metatarsal organ culture reproducing bone collar formation , activation of BMP signaling enhanced the bone collar formation cooperatively with Hh input , whereas the signaling induced ectopic chondrocyte formation in the perichondrium without Hh input .
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Please list all entity words in the text that fit the category.Output format is "type1: word1; type2: word2". 
Option: Chemical 
Text: {0} 
Answer:
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