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Particle bombardment of barley aleurone with a B22EL8 promoter - GUS ( beta - glucuronidase ) construct demonstrates that the promoter ( 3 kb ) is active in developing barley grains .
 GENE: B22EL8 promoter; GENE: GUS; GENE: beta - glucuronidase
Please list all entity words in the text that fit the category.Output format is "type1: word1; type2: word2". 
Option: GENE 
Text: {0} 
Answer:
 GENE: B22EL8 promoter; GENE: GUS; GENE: beta - glucuronidase