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Using a combination of phylogeographic analyses and circuit theory , I investigate how characteristics of landscape heterogeneity including regional distributions of slope , rivers and streams , habitat and hydrological basins ( drainages ) impact genetic distance among populations of the endemic spotted reed frog ( Hyperolius substriatus ) , identifying corridors of connectivity as well as barriers to dispersal .
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Please list all entity words in the text that fit the category.Output format is "type1: word1; type2: word2". 
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