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Reversing the position and subsequent mutagenesis of the L1 and L2 domains indicated that the primary sequence of these motifs rather than their position in the E3 promoter was critical for regulating gene expression .
 DNA: primary sequence; DNA: L1; DNA: E3 promoter; DNA: L2; DNA: L2 domains
Please list all entity words in the text that fit the category.Output format is "type1: word1; type2: word2". 
Option: cell line, protein, RNA, DNA, cell type 
Text: {0} 
Answer:
 DNA: primary sequence; DNA: L1; DNA: E3 promoter; DNA: L2; DNA: L2 domains