1039
Since the end of the Pleistocene , the three - spined stickleback ( Gasterosteus aculeatus ) has repeatedly colonized and adapted to various freshwater habitats probably originating from ancestral marine populations .
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Please list all entity words in the text that fit the category.Output format is "type1: word1; type2: word2". 
Option: Chemical 
Text: {0} 
Answer:
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