1840
We also show that VEGFR1 + cells express VLA - 4 ( also known as integrin alpha4beta1 ) , and that tumour - specific growth factors upregulate fibronectin - - a VLA - 4 ligand - - in resident fibroblasts , providing a permissive niche for incoming tumour cells .
 Anatomy: VEGFR1 + cells; Anatomy: tumour; Anatomy: fibroblasts; Anatomy: tumour cells
Please list all entity words in the text that fit the category.Output format is "type1: word1; type2: word2". 
Option: Anatomy 
Text: {0} 
Answer:
 Anatomy: VEGFR1 + cells; Anatomy: tumour; Anatomy: fibroblasts; Anatomy: tumour cells