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Our results collectively demonstrate that overdosage of Hand2 is a major cause for at least the limb and heart phenotypes of 4q + and that mouse Rim4 provides a unique animal model for understanding the molecular bases underlying the complex phenotypes of 4q + .
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Please list all entity words in the text that fit the category.Output format is "type1: word1; type2: word2". 
Option: Chemical 
Text: {0} 
Answer:
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