3903
While Piccolo NuA4 ' s catalytic subunit Esa1 alone is unable to acetylate nucleosomal histones , its accessory subunits , Yng2 and Epl1 , enable Esa1 to bind to and to act on nucleosomes .
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Please list all entity words in the text that fit the category.Output format is "type1: word1; type2: word2". 
Option: Chemical 
Text: {0} 
Answer:
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