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Disruption of INP53 , but not the related INP51 and INP52 genes , resulted in alpha - factor maturation defects and exacerbated alpha - factor maturation defects when combined with chc1 - 521 .
 GENE: INP53; GENE: INP51; GENE: INP52 genes; GENE: alpha - factor; GENE: alpha - factor; GENE: chc1 - 521
Please list all entity words in the text that fit the category.Output format is "type1: word1; type2: word2". 
Option: GENE 
Text: {0} 
Answer:
 GENE: INP53; GENE: INP51; GENE: INP52 genes; GENE: alpha - factor; GENE: alpha - factor; GENE: chc1 - 521