1673
The results indicate that expression of proU in E . coli is directed from two promoters , one ( P2 ) characterized earlier by other workers with the start site of transcription 60 nucleotides upstream of the initiation codon of the first structural gene ( proV ) , and the other ( P1 ) situated 250 nucleotides upstream of proV .
 GENE: proU; GENE: proV; GENE: proV
Please list all entity words in the text that fit the category.Output format is "type1: word1; type2: word2". 
Option: GENE 
Text: {0} 
Answer:
 GENE: proU; GENE: proV; GENE: proV