4164
During chordotonal organ development , the 3 ' enhancer directs expression in proneural clusters ; whereas successive modular enhancers located in the 5 ' region drive tissue - specific expression in chordotonal organ precursors in the embryo and larval leg , wing and antennal imaginal discs .
 None
Please list all entity words in the text that fit the category.Output format is "type1: word1; type2: word2". 
Option: GENE 
Text: {0} 
Answer:
 None