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Selection on quantitative trait loci ( QTL ) may vary among natural environments due to differences in the genetic architecture of traits , environment - specific allelic effects or changes in the direction and magnitude of selection on specific traits .
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Given options, please tell me the categories of all the listed entity words.Output format is "type1: word1; type2: word2". 
Option: Chemical 
Text: {0} 
 Entities:  
Answer:
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