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Cotransfection experiments suggest that HNF - 3beta , HNF - 1alpha , and SP1 are positive regulators of the herein - described human PDX - 1 enhancer element .
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Please list all entity words in the text that fit the category.Output format is "type1: word1; type2: word2". 
Option: Anatomy 
Text: {0} 
Answer:
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