1669
TSA treatment , however , did not detectably alter enhancer factor binding or the positioning of nuc - 1 on the majority of the chromatin templates indicating that protein acetylation and chromatin remodeling may be limiting steps that occur only on transcriptionally competent templates , or that remodeling of nuc - 1 requires additional factors .
 GENE: nuc - 1; GENE: nuc - 1
Please list all entity words in the text that fit the category.Output format is "type1: word1; type2: word2". 
Option: GENE 
Text: {0} 
Answer:
 GENE: nuc - 1; GENE: nuc - 1