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DNase I footprinting analysis revealed two protected regions , one binding the transcription factors SP1 and SP3 and the other hepatocyte nuclear factor 3beta ( HNF - 3beta ) and HNF - 1alpha .
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Please list all entity words in the text that fit the category.Output format is "type1: word1; type2: word2". 
Option: Anatomy 
Text: {0} 
Answer:
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