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Several QTL were pleiotropic across environments , three colocalizing with known functional polymorphisms in flowering time genes ( CRY2 , FRI and MAF2 - 5 ) , but major QTL differed across field sites , showing conditional neutrality .
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Please list all entity words in the text that fit the category.Output format is "type1: word1; type2: word2". 
Option: Chemical 
Text: {0} 
Answer:
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