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In two oceanic field sites with high prereproductive mortality , QTL alleles contributing to early reproduction resulted in greater fruit production , conferring selective advantage , whereas alleles contributing to later reproduction resulted in larger size and higher fitness in a continental site .
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Please list all entity words in the text that fit the category.Output format is "type1: word1; type2: word2". 
Option: Chemical 
Text: {0} 
Answer:
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