1988
Two of the mutations included a G to C substitution at codon 12 , and a G to A substitution at codon 18 , resulting in a glycine to arginine , and an alanine to threonine change at these amino acids , respectively .
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Please list all entity words in the text that fit the category.Output format is "type1: word1; type2: word2". 
Option: Anatomy 
Text: {0} 
Answer:
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