1394
Transcription initiation sites of the rat II beta - , III beta , and O beta - globin genes were determined to be 52 base pairs ( bp ) 5 ' - upstream of the translation initiation codon ( ATG ) , in each gene by primer extension analysis .
 GENE: II beta - , III beta , and O beta - globin
Please list all entity words in the text that fit the category.Output format is "type1: word1; type2: word2". 
Option: GENE 
Text: {0} 
Answer:
 GENE: II beta - , III beta , and O beta - globin