1175
One complex most likely contained Sp1 , and another complex showed S - phase - specific binding , suggesting a direct role in the cell - cycle - dependent R1 gene expression .
 GENE: Sp1; GENE: R1 gene
Please list all entity words in the text that fit the category.Output format is "type1: word1; type2: word2". 
Option: GENE 
Text: {0} 
Answer:
 GENE: Sp1; GENE: R1 gene