6111
Furthermore , the negative transcriptional effect of COUP - TFs is dominant over the activating effect of the Oct4 embryonic stem cell - specific enhancer .
 COUP - TFs, Oct4 embryonic stem cell - specific enhancer
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: GENE 
Text: {0} 
Answer:
 COUP - TFs, Oct4 embryonic stem cell - specific enhancer