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Gel - mobility - shift assays confirmed that the Raji nuclear proteins that bound to W and V elements were competed with by an HLA - DRA X - box oligonucleotide .
 HLA - DRA X - box oligonucleotide
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: GENE 
Text: {0} 
Answer:
 HLA - DRA X - box oligonucleotide