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It attenuates the signaling activity of G - proteins , blocking the homing of Intra Epithelial Lymphocytes ( IELs ) , and it is specifically expressed both in human small intestinal mucosa and in murine IELs , key players in the development of human CD villous atrophy [ 7 ] , [ 17 ] .
 Intra Epithelial Lymphocytes, IELs, small intestinal mucosa, IELs
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: Anatomy 
Text: {0} 
Answer:
 Intra Epithelial Lymphocytes, IELs, small intestinal mucosa, IELs