1159
Using avidin - biotin complex DNA binding assays , a series of overlapping alpha promoter DNA sequences between - 170 to 29 basepairs were tested , but each failed to bind GR , whereas a control GRE avidly bound receptor .
 avidin - biotin, alpha promoter DNA sequences, GR
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: GENE 
Text: {0} 
Answer:
 avidin - biotin, alpha promoter DNA sequences, GR