6502
These cell lines displayed methylation of the CpG island surrounding the first exon of p16INK4A and expressed abundant levels of a nontranslated mRNA containing an alternative first exon ( E1 beta ) , as did all other cell lines in which the p16INK4A locus was not deleted .
 p16INK4A, p16INK4A locus
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: GENE 
Text: {0} 
Answer:
 p16INK4A, p16INK4A locus