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" Synodontis serratus " , known as the " shield-head squeaker " , is a species of upside-down catfish that is native to the Nile basin of the Democratic Republic of the Congo , Egypt , Ethiopia and Sudan .
 Nile, Democratic Republic of the Congo, Egypt, Ethiopia, Sudan
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: media, plant, vehicle, astronomical object, time, disease, animal, location, person, organization, mythical figure, food, biology, geographical phenomenon, instrument, event 
Text: {0} 
Answer:
 Nile, Democratic Republic of the Congo, Egypt, Ethiopia, Sudan