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The composition of these forests varies throughout their range but often includes chestnut oak ( " Quercus prinus " ) , northern red oak ( " Quercus rubra " ) , eastern black oak ( " Quercus velutina " ) , white oak , ( " Quercus alba " ) , and scarlet oak ( " Quercus coccinea " ) .
 Quercus rubra, Quercus velutina, Quercus alba, Quercus coccinea
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: media, plant, vehicle, astronomical object, time, disease, animal, location, person, organization, mythical figure, food, biology, geographical phenomenon, instrument, event 
Text: {0} 
Answer:
 Quercus rubra, Quercus velutina, Quercus alba, Quercus coccinea