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Interestingly , the positions of these introns have been conserved in comparison with the genes of two other transglutaminase - like activities described in the literature , but the TGM1 gene is by far the smallest characterized to date because its introns are relatively smaller .
 TGM1 gene
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: GENE 
Text: {0} 
Answer:
 TGM1 gene