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He has authored taxonomic descriptions of 140 species , subspecies and varieties of plants , especially in the palm family Henderson was educated in Wycliffe College in Gloucestershire and Birkbeck College , University of London , and received his Ph.D. from City University of New York in 1987 .
 Henderson, Wycliffe College, Gloucestershire, Birkbeck College , University of London, City University of New York
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: location, person, organization 
Text: {0} 
Answer:
 Henderson, Wycliffe College, Gloucestershire, Birkbeck College , University of London, City University of New York