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The granulocyte-macrophage colony-stimulating factor ( GM-CSF ) gene has been studied extensively as a model system of transcriptional induction during T-lymphocyte activation .
 granulocyte-macrophage colony-stimulating factor ( GM-CSF ) gene, granulocyte-macrophage colony-stimulating factor, GM-CSF
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: cell line, protein, RNA, DNA, cell type 
Text: {0} 
Answer:
 granulocyte-macrophage colony-stimulating factor ( GM-CSF ) gene, granulocyte-macrophage colony-stimulating factor, GM-CSF