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p130 , predominantly in Form 1, and hypophosphorylated pRb bind an E2F DNA binding site ; p130 complexes E2F-4 , whereas pRb binds both E2F-4 and E2F-1 .
 E2F-4, E2F DNA binding site, p130, pRb, hypophosphorylated pRb, E2F-1, p130, pRb
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: cell line, protein, RNA, DNA, cell type 
Text: {0} 
Answer:
 E2F-4, E2F DNA binding site, p130, pRb, hypophosphorylated pRb, E2F-1, p130, pRb