9107
However , while IL - 1beta and TNF - alpha both induced nuclear binding of the Rel proteins p50 and p65 to an NF - kappaB consensus oligonucleotide in gel shift assays and caused transient degradation of inhibitor of NF - kappaB - alpha ( IkappaB - alpha ) in the cytoplasm of myofibroblasts , only IL - 1beta upregulated PDGF - Ralpha .
 IL - 1beta, TNF - alpha, Rel proteins, p50, p65, NF - kappaB consensus oligonucleotide, inhibitor of NF - kappaB - alpha, IkappaB - alpha, IL - 1beta, PDGF - Ralpha
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: GENE 
Text: {0} 
Answer:
 IL - 1beta, TNF - alpha, Rel proteins, p50, p65, NF - kappaB consensus oligonucleotide, inhibitor of NF - kappaB - alpha, IkappaB - alpha, IL - 1beta, PDGF - Ralpha