8608
Chimaeric VP16 - E2 molecules suggest that the epithelial specific transcriptional activation of the BPV - 4 LCR promoter is mediated by the E2 transactivation domain .
 VP16, E2, BPV - 4 LCR promoter, E2 transactivation domain
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: GENE 
Text: {0} 
Answer:
 VP16, E2, BPV - 4 LCR promoter, E2 transactivation domain