1272
A 330 - kb YAC , A148A7 , spanned from the HLA - DQA1 locus through the Y3 / Ring 4 locus and extended at least 130 kb centromeric of YAC B1D12 .
 HLA, DQA1, Y3, Ring 4
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: GENE 
Text: {0} 
Answer:
 HLA, DQA1, Y3, Ring 4