11148
Promoter activity was dose - dependently inhibited by cotransfection with either ras or mos oncogenes , but oncogene inhibition was reversed and the overall activity increased when cells were treated with the MAP kinase kinase ( MKK ) inhibitor PD98059 .
 ras, mos oncogenes, MAP kinase kinase, MKK
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: GENE 
Text: {0} 
Answer:
 ras, mos oncogenes, MAP kinase kinase, MKK