5770
Mutagenesis of each NRF - 1 motif in the ALAS promoter gave substantially lowered levels of chloramphenicol acetyltransferase expression , whereas mutagenesis of both NRF - 1 motifs resulted in the almost complete loss of expression .
 NRF - 1 motif, ALAS promoter, chloramphenicol acetyltransferase, NRF - 1 motifs
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: GENE 
Text: {0} 
Answer:
 NRF - 1 motif, ALAS promoter, chloramphenicol acetyltransferase, NRF - 1 motifs