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The Paraguay River flows through the Pantanal wetland , after which its main tributaries include the Pilcomayo River and the Bermejo River , before it ends in the Paraná .
 Paraguay River, Pantanal, Pilcomayo River, Bermejo River, Paraná
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: location, person, organization 
Text: {0} 
Answer:
 Paraguay River, Pantanal, Pilcomayo River, Bermejo River, Paraná