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Protein-DNA binding studies suggested that Sp1 and two other CCGCCC binding proteins from erythroid and non-erythroid cells could bind to the Sp1 binding motif .
 Sp1 binding motif, CCGCCC binding proteins, Sp1, Sp1
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: cell line, protein, RNA, DNA, cell type 
Text: {0} 
Answer:
 Sp1 binding motif, CCGCCC binding proteins, Sp1, Sp1