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Nuclei from GATA-1 -mutant embryonic stem cells can still be reprogrammed to express their globin genes in erythroid heterokaryons , indicating that de novo induction of GATA-1 is not required for globin gene activation following cell fusion .
 GATA-1 -mutant embryonic stem cells, erythroid heterokaryons, embryonic stem cells, globin genes, GATA-1, GATA-1, globin gene
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: cell line, protein, RNA, DNA, cell type 
Text: {0} 
Answer:
 GATA-1 -mutant embryonic stem cells, erythroid heterokaryons, embryonic stem cells, globin genes, GATA-1, GATA-1, globin gene