11012
Inbreeding in R . flavipes was most strongly associated with climatic variables ( mean annual temperature and seasonality ) , whereas nonclimatic variables , including the availability of wood substrate and soil composition , were important predictors of inbreeding in R . grassei .
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Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: Chemical 
Text: {0} 
Answer:
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