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By multimerizing the regulatory elements of the GM-CSF promoter , we found that one of the target sites for the CN action is the conserved lymphokine element 0 ( CLE0 ), located at positions between -54 and -40 .
 GM-CSF, GM-CSF promoter, -54 and -40, CN, CLE0, regulatory elements, target sites, conserved lymphokine element 0
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: cell line, protein, RNA, DNA, cell type 
Text: {0} 
Answer:
 GM-CSF, GM-CSF promoter, -54 and -40, CN, CLE0, regulatory elements, target sites, conserved lymphokine element 0