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We found that expression of a constitutively active form of calcineurin could substitute for Ca2+ ionophore and synergize with PMA to activate the GM-CSF promoter , and conversely that mutant-activated Ras could substitute for PMA and cooperate with Ca2+ ionophore .
 GM-CSF promoter, calcineurin, GM-CSF, mutant-activated Ras
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: cell line, protein, RNA, DNA, cell type 
Text: {0} 
Answer:
 GM-CSF promoter, calcineurin, GM-CSF, mutant-activated Ras