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The combination of sophisticated wet-rice agriculture , based on an engineered irrigation system and the Tonlé Sap 's spectacular abundance in fish and aquatic fauna , as protein source guaranteed a regular food surplus .
 Tonlé Sap
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: media, plant, vehicle, astronomical object, time, disease, animal, location, person, organization, mythical figure, food, biology, geographical phenomenon, instrument, event 
Text: {0} 
Answer:
 Tonlé Sap