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Together , these results indicate that IL - 1 beta induces VEGF gene expression at both transcriptional and post - transcriptional levels , and IL - 1 beta evokes p38 MAPK and JNK signalings , which in turn stimulate the transcription of the VEGF gene through Sp1 - binding sites .
 IL - 1 beta, VEGF gene, IL - 1 beta, p38, JNK, VEGF gene, Sp1 - binding sites
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: GENE 
Text: {0} 
Answer:
 IL - 1 beta, VEGF gene, IL - 1 beta, p38, JNK, VEGF gene, Sp1 - binding sites