17785
" Metaloricaria " is only known from the Guiana Shield in French Guiana and Suriname where the species occupy an ecological niche similar to that of " Harttia " .
 Guiana Shield, French Guiana, Suriname, Harttia
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: media, plant, vehicle, astronomical object, time, disease, animal, location, person, organization, mythical figure, food, biology, geographical phenomenon, instrument, event 
Text: {0} 
Answer:
 Guiana Shield, French Guiana, Suriname, Harttia