23758
Pheromones from species of Rutelinae are produced from epithelial cells lining the inner surface of the apical abdominal segments ; amino acid-based pheromones of Melolonthinae are produced from eversible glands on the abdominal apex .
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Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: location, person, organization 
Text: {0} 
Answer:
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