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Finally , profiling of a gcn4Delta mutant uncovered an alternative induction pathway operating at many Gcn4p target genes in histidine - starved cells .
 gcn4Delta mutant, Gcn4p target genes
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: GENE 
Text: {0} 
Answer:
 gcn4Delta mutant, Gcn4p target genes