98159
In 1921 , the evolutionary biologist Richard Goldschmidt argued that the observed increase in the melanic form of the black arches moth , " Lymantria monacha " , could not have been caused by mutation pressure alone , but required a selective advantage from an unknown cause : he did not consider camouflage as an explanation .
 Richard Goldschmidt
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: media, plant, vehicle, astronomical object, time, disease, animal, location, person, organization, mythical figure, food, biology, geographical phenomenon, instrument, event 
Text: {0} 
Answer:
 Richard Goldschmidt