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We investigated the role of this G + C-rich region in augmenting alpha-globin promoter activity in the presence of the far-upstream alpha-globin enhancer , HS-40 .
 HS-40, G + C-rich region, alpha-globin promoter, far-upstream alpha-globin enhancer
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: cell line, protein, RNA, DNA, cell type 
Text: {0} 
Answer:
 HS-40, G + C-rich region, alpha-globin promoter, far-upstream alpha-globin enhancer