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In the 19th century , particularly after the publication of " On the Origin of Species " in 1859 , the idea that life had evolved was an active source of academic debate centred on the philosophical , social and religious implications of evolution .
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Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: location, person, organization 
Text: {0} 
Answer:
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