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When the promoter region was linked with a heterologous reporter gene , we found that the promoter region is inducible by both interferons ( interferon -alpha and -gamma ) and interferon regulatory factor 1 .
 promoter region, heterologous reporter gene, promoter region, interferon regulatory factor 1, interferons
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: cell line, protein, RNA, DNA, cell type 
Text: {0} 
Answer:
 promoter region, heterologous reporter gene, promoter region, interferon regulatory factor 1, interferons