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A novel SP-1 site in the human interleukin-1 beta promoter confers preferential transcriptional activity in keratinocytes .
 human interleukin-1 beta promoter, novel SP-1 site, keratinocytes, SP-1 site
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: cell line, protein, RNA, DNA, cell type 
Text: {0} 
Answer:
 human interleukin-1 beta promoter, novel SP-1 site, keratinocytes, SP-1 site