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Typical woody understory and shrub layer species include moosewood ( " Acer pensylvanicum " ) , low-bush blueberry ( " Vaccinium angustifolium " ) and other heath species , especially the genera " Gaylussacia " and " Vaccinium " .
 Acer pensylvanicum, Vaccinium angustifolium, Gaylussacia, Vaccinium
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
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Answer:
 Acer pensylvanicum, Vaccinium angustifolium, Gaylussacia, Vaccinium