1586
The gcd2 - 503 mutation also results in polysome runoff , accumulation of inactive 80S ribosomal couples , and accumulation of at least one of the subunits of the general translation initiation factor 2 ( eIF - 2 alpha ) in 43S - 48S particles following a shift to the restrictive temperature .
 gcd2 - 503, 80S ribosomal couples, general translation initiation factor 2, eIF - 2 alpha, 43S, 48S
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: GENE 
Text: {0} 
Answer:
 gcd2 - 503, 80S ribosomal couples, general translation initiation factor 2, eIF - 2 alpha, 43S, 48S