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The two related protein- tyrosine kinases Syk and Zap are rapidly phosphorylated on tyrosine residues and enzymatically activated upon crosslinking of the T cell antigen receptor .
 T cell antigen receptor, Zap, Syk, protein- tyrosine kinases
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: cell line, protein, RNA, DNA, cell type 
Text: {0} 
Answer:
 T cell antigen receptor, Zap, Syk, protein- tyrosine kinases