12784
NBH embryos had lower basal expression of antioxidant genes catalase and glutathione - S - transferase alpha ( gsta ) , and upon exposure to tBHQ , exhibited lower levels of expression of catalase , gsta , and superoxide dismutase compared to controls .
 glutathione, S, tBHQ, superoxide
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: Chemical 
Text: {0} 
Answer:
 glutathione, S, tBHQ, superoxide