13504
The locus control region of the beta-globin cluster contains five DNase I hypersensitive sites ( 5'HS1-5 ) required for locus activation .
 beta-globin cluster, 5'HS1-5, DNase I hypersensitive sites, locus control region
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: cell line, protein, RNA, DNA, cell type 
Text: {0} 
Answer:
 beta-globin cluster, 5'HS1-5, DNase I hypersensitive sites, locus control region