110949
Preferential gene retention after a genome-wide duplication event in the genus " Nicotiana " partially accounts for this large size , which is roughly twice that of " N. obtusifolia " ( ~ 1.23 Gb ) , a closely related species .
 Nicotiana, N. obtusifolia
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: media, plant, vehicle, astronomical object, time, disease, animal, location, person, organization, mythical figure, food, biology, geographical phenomenon, instrument, event 
Text: {0} 
Answer:
 Nicotiana, N. obtusifolia