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In 1927 , Nikolai Koltsov proposed that inherited traits would be inherited via a " giant hereditary molecule " made up of " two mirror strands that would replicate in a semi-conservative fashion using each strand as a template " .
 Nikolai Koltsov
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: location, person, organization 
Text: {0} 
Answer:
 Nikolai Koltsov