80833
The rate of release is lowest in the shelled cephalopods " Nautilus " and " Sepia " as a result of their using nitrogen to fill their shells with gas to increase buoyancy .
 Nautilus, Sepia
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: media, plant, vehicle, astronomical object, time, disease, animal, location, person, organization, mythical figure, food, biology, geographical phenomenon, instrument, event 
Text: {0} 
Answer:
 Nautilus, Sepia