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Here we present evidence that major IE proteins IE86 , IE72 , and IE55 are capable of trans - activating the HIV LTR in a T - cell line , HUT - 78 .
 major IE proteins, IE86, IE72, IE55, HIV LTR
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: GENE 
Text: {0} 
Answer:
 major IE proteins, IE86, IE72, IE55, HIV LTR