110888
However , when FOXO1 is phosphorylated by Akt on Thr-24 , Ser-256 , and Ser-319 , it is excluded from the nucleus , where it is then ubiquitinated and degraded .
 None
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: location, person, organization 
Text: {0} 
Answer:
 None