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Primer extension and ribonuclease protection analyses indicate a single major transcriptional start site .
 major transcriptional start site, ribonuclease
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: cell line, protein, RNA, DNA, cell type 
Text: {0} 
Answer:
 major transcriptional start site, ribonuclease