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The major putative downstream site for E2F , located in the first exon , serves as a target for E2F-1/DP1 complex binding in vitro.
 major putative downstream site, E2F, E2F-1/DP1 complex, first exon
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: cell line, protein, RNA, DNA, cell type 
Text: {0} 
Answer:
 major putative downstream site, E2F, E2F-1/DP1 complex, first exon