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These studies indicate that maximal LPS induction of the TNF-alpha promoter is mediated by concerted participation of at least two separate cis-acting regulatory elements .
 TNF-alpha, cis-acting regulatory elements, TNF-alpha promoter
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: cell line, protein, RNA, DNA, cell type 
Text: {0} 
Answer:
 TNF-alpha, cis-acting regulatory elements, TNF-alpha promoter