2869
Contrary to the observation made with other S . pombe genes transcribed in the budding yeast , the heterologous actin gene transcript is initiated 39 nucleotides upstream of the initiation start site used in the homologous yeast .
 actin gene transcript
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: GENE 
Text: {0} 
Answer:
 actin gene transcript