8982
In contrast , transcription initiating from opuE P - 1 ( sigma A ) rose in proportion to the external osmolarity and was maintained at high levels .
 opuE P - 1, sigma A
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: GENE 
Text: {0} 
Answer:
 opuE P - 1, sigma A