65401
Higher temperatures during the Miocene or sea level oscillations may have separated early populations , leading to allopatric speciation , although admixture and introgression between lineages probably took place .
 Miocene
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: location, person, organization 
Text: {0} 
Answer:
 Miocene