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A 2012 study using 18 motion-sensitive cameras in Río Los Cipreses National Reserve counted a population of two males and two females , one of them with at least two cubs , in an area of 600 " km ( 0.63 cougars per 100 " km ) .
 Río Los Cipreses National Reserve
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: media, plant, vehicle, astronomical object, time, disease, animal, location, person, organization, mythical figure, food, biology, geographical phenomenon, instrument, event 
Text: {0} 
Answer:
 Río Los Cipreses National Reserve