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Monocyte-macrophage differentiation was used as a model system for studying gene regulation of the human vacuolar H(+)-ATPase ( V-ATPase ).
 V-ATPase, human vacuolar H(+)-ATPase
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: cell line, protein, RNA, DNA, cell type 
Text: {0} 
Answer:
 V-ATPase, human vacuolar H(+)-ATPase