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Infection with Theileria annulata induces expression of matrix metalloproteinase 9 and transcription factor AP-1 in bovine leucocytes .
 AP-1, transcription factor, matrix metalloproteinase 9, bovine leucocytes
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: cell line, protein, RNA, DNA, cell type 
Text: {0} 
Answer:
 AP-1, transcription factor, matrix metalloproteinase 9, bovine leucocytes