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The upregulated species of AE2 mRNA originates from the classical upstream promoter of the AE2 gene ( here referred to as AE2a1 ) which provides the binding site for transcription factors early growth response 1 ( EGR1 ) and SP1 .
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Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: Chemical 
Text: {0} 
Answer:
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