47286
Research using mitochondrial rRNA suggests that these three species descended from a common ancestor , and that they are more closely related with " Polistes nimpha " and " Polistes dominula " than with " Polistes gallicus " and " Polistes biglumis " .
 Polistes gallicus, Polistes biglumis
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: media, plant, vehicle, astronomical object, time, disease, animal, location, person, organization, mythical figure, food, biology, geographical phenomenon, instrument, event 
Text: {0} 
Answer:
 Polistes gallicus, Polistes biglumis