8618
The COOH - terminus of this new isoform , which we designate beta 4 , lacks a 22 amino acid lysine - rich sequence common to both the human red cell alpha - and beta - adducin subunits and homologous to a highly conserved region in MARCKS , a filamentous actin - cross linking protein regulated by protein kinase C and calcium / calmodulin . beta 4 - adducin preserves a previously identified calmodulin binding domain .
 beta 4, human red cell alpha - and beta - adducin subunits, MARCKS, actin, protein kinase C, calmodulin, beta 4 - adducin, calmodulin binding domain
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: GENE 
Text: {0} 
Answer:
 beta 4, human red cell alpha - and beta - adducin subunits, MARCKS, actin, protein kinase C, calmodulin, beta 4 - adducin, calmodulin binding domain