5458
Cytoplasmic dynein is a multisubunit , microtubule - dependent mechanochemical enzyme that has been proposed to function in a variety of intracellular movements , including minus - end - directed transport of organelles .
 Cytoplasmic dynein
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: GENE 
Text: {0} 
Answer:
 Cytoplasmic dynein