11681
In strains lacking tup11 and tup12 , the atf1 - pcr1 transcriptional activator continues to play a central role in fbp1 - lacZ expression ; however , spc1 MAPK phosphorylation of atf1 is no longer essential for its activation .
 tup11, tup12, atf1, pcr1, fbp1, lacZ, spc1 MAPK, atf1
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: GENE 
Text: {0} 
Answer:
 tup11, tup12, atf1, pcr1, fbp1, lacZ, spc1 MAPK, atf1