9969
Site-directed mutational studies demonstrate that Site I and Site IV together support ATF1- and CREB- induced trans-activation of the H4 promoter .
 Site I, Site IV, H4 promoter
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: cell line, protein, RNA, DNA, cell type 
Text: {0} 
Answer:
 Site I, Site IV, H4 promoter