25136
Whereas Lys165 and Arg33 were required for efficient catalysis or phosphosubstrate binding of PPM1A , Gln185 and Arg186 were determinants of PPM1A substrate specificity .
 Lys165, Arg33, Gln185, Arg186
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: Chemical 
Text: {0} 
Answer:
 Lys165, Arg33, Gln185, Arg186