2667
In addition to p16 ( INK4A ) , the INK4A locus encodes a splice variant , termed p12 - - specifically expressed in human pancreas - - and ARF , a protein encoded by an alternative reading frame that acts as a tumor suppressor through the p53 pathway .
 pancreas, tumor
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: Anatomy 
Text: {0} 
Answer:
 pancreas, tumor