5391
Our results suggest that the bulk of protein-DNA interactions that underlies the developmental control of globin genes takes place in the gamma- and beta- globin gene promoters , and that GT motifs of the beta-globin locus LCR may play a role in the developmental regulation of human beta-globin gene expression , perhaps by increasing the probability of interaction of the LCR holocomplex with the fetal or the adult globin gene .
 LCR, beta-globin locus, LCR holocomplex, LCR, GT motifs, beta-globin gene, globin genes
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: cell line, protein, RNA, DNA, cell type 
Text: {0} 
Answer:
 LCR, beta-globin locus, LCR holocomplex, LCR, GT motifs, beta-globin gene, globin genes