8898
However , mutations in the HNF - 4 binding site on element CIIB and inhibition of HNF - 4 synthesis in HepG2 cells by antisense HNF - 4 constructs decreased the apoCII promoter activity to 25 - 40 % of the control , indicating that HNF - 4 is a positive regulator of the apoCII gene .
 HNF - 4 binding site, element CIIB, HNF - 4, HNF - 4 constructs, apoCII promoter, HNF - 4, apoCII gene
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: GENE 
Text: {0} 
Answer:
 HNF - 4 binding site, element CIIB, HNF - 4, HNF - 4 constructs, apoCII promoter, HNF - 4, apoCII gene