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His work contributed to the conceptual revolution that led to the modern evolutionary synthesis of Mendelian genetics , systematics , and Darwinian evolution , and to the development of the biological species concept .
 Mendelian, Darwinian
Please list all entity words in the text that fit the category. Output format is "word1, word2". 
Option: location, person, organization 
Text: {0} 
Answer:
 Mendelian, Darwinian