When he crossed varieties which differed in a trait (e.g. tall crossed with short), the first generation of hybrids (F1) showed only one of the two alternatives. One character was "dominant", and the other "recessive". But when he crossed these hybrids with each other, the recessive character reappeared in the second (F2) generation. The proportion of plants showing the dominant as opposed to the recessive character was close to 3 to 1. Further analysis of the descendants (F3) of the dominant group showed that one-third of them were true-breeding and two-thirds were of hybrid constitution. The 3:1 ratio could therefore be rewritten as 1:2:1, meaning that 50 percent of the F2 generation were true-breeding and 50 percent were still hybrid. This was Mendel’s major discovery.