Therefore , physiologist Hugh Davson and biologist James Danielli suggested that membranes indeed do have proteins .
For instance , their model could not provide answers to questions on surface tension , permeability , and the electric resistance of membranes .
Physiologist Hugh Davson and biologist James Danielli suggested that membranes indeed do have proteins .
For instance , their model could not provide answers to questions on surface tension , permeability , and the electric resistance of membranes .