Travelling to the UK on a Shell Overseas Science Scholarship , he performed his PhD work at University College London under the supervision of Franz Sondheimer .
Holmes ' undergraduate studies and masters ' research were conducted at the University of Melbourne in Australia .
Travelling to the UK on a Shell Overseas Science Scholarship , Holmes performed Holmes ' PhD work at University College London under the supervision of Franz Sondheimer .
Holmes ' undergraduate studies and masters ' research were conducted at the University of Melbourne in Australia .