He lost his right hand at 40 years of age , and began painting with left hand .
He was the successor of Knoller as teacher of painting at the Brera Academy as well as conservator of the Pinacoteca .
successor lost successor 's right hand at 40 years of age , and began painting with left hand .
He was the successor of Knoller as teacher of painting at the Brera Academy as well as conservator of the Pinacoteca .