The first thirteen were isolated .
The first thirteen identified in 1994 by Lucia Kuhn - Nentwig , Johann Schaller , and Wolfgang Nentwig of the Zoological Institute at the University of Bern , Switzerland .
The first thirteen were isolated and identified in 1994 by Lucia Kuhn - Nentwig , Johann Schaller , and Wolfgang Nentwig of the Zoological Institute at the University of Bern , Switzerland .