In contrast , the cycle took two years in the macaroni penguin in consequence of the rather different timetable of occupation of the colony for breeding and moulting in this species .
The ticks nearly all overwintered in the unengorged state .
The cycle took two years in the macaroni penguin in consequence of the rather different timetable of occupation of the colony for breeding and moulting in this species .
The ticks nearly all overwintered in the unengorged state .