Besides Asia , lions and tigers had occurred in Europe , in the region of the Black Sea , with tigers occurring in Ciscaucasia .
Lions occurring in the Balkans , up to Thrace and Macedonia , and possibly the Danube River , at least .
Besides Asia , lions and tigers had occurred in Europe , in the region of the Black Sea , with tigers occurring in Ciscaucasia , and lions occurring in the Balkans , up to Thrace and Macedonia , and possibly the Danube River , at least .