PERSON@1 -LRB- April NUMBER@1 , NUMBER@2 - November NUMBER@3 , NUMBER@4 -RRB- was an MISC@1 geneticist and physician whose most significant contribution to present-day biology was his theory that the MISC@2 laws of inheritance could be applied to chromosomes at the cellular level of living organisms .

PERSON@1 -LRB- April NUMBER@1 , NUMBER@2 - November NUMBER@3 , NUMBER@4 -RRB- was an MISC@1 biologist .
