PERSON@1 -LRB- -RRB- -LRB- also romanized PERSON@2 -RRB- -LRB- NUMBER@1 May NUMBER@2 -- NUMBER@3 July NUMBER@4 -RRB- was a MISC@1 microbiologist best remembered for his pioneering research into the immune system .

PERSON@1 -LRB- May NUMBER@1 , NUMBER@2 - July NUMBER@1 , NUMBER@4 -RRB- was a MISC@1 biologist .
