Proceedings of the ORGANIZATION@1 NUMBER@1 : NUMBER@2 In this case , an individual 's inclusive fitness varies depending on how much it invests in within-group competition -LRB- e.g. hoarding a private food cache -RRB- versus between-group competition -LRB- e.g. contributing to common foraging -RRB- ; and on its relatedness to the other group members .

In their model , an individual 's inclusive fitness varies depending on how much it invests in within-group competition -LRB- e.g. hoarding a private food cache -RRB- versus between-group competition -LRB- e.g. contributing to common foraging -RRB- ; and on its relatedness to the other group members .
