Begun says that these primates flourished in LOCATION@1 and that the lineage leading to the MISC@1 apes and humans -- including Dryopithecus -- migrated south from LOCATION@2 or LOCATION@3 into LOCATION@4 .

Begun says that these primates flourished in LOCATION@1 and that their lineage leading to the MISC@1 apes and humans -- Dryopithecus -- migrated south from LOCATION@2 or LOCATION@3 into LOCATION@4 .
