While groups may benefit from avoiding certain behaviors, those harmful behaviors have the same effect regardless of whether the offending individuals are aware of them or not. Since the individuals themselves can increase their reproductive success by doing many of them, any characteristics that entail impunity are positively selected by evolution. Specifically punishing individuals aware of their breach of rules would select against the ability to be aware of it, precluding any coevolution of both conscious choice and a sense of it being the basis for moral and penal liability in the same species.

শাস্তি সমস্যা (The punishment problems)
সু
ধর্মের বিবর্তন
প্রাইমেট সামাজিকতা (Primate sociality)