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tokens: Acknowledgements We thank Alexandre Faure for providing collisional excitation rates at low temperatures, and Luca Matrà for sharing the model of the dust radiation field and the ALMA data of CO. We also thank the referee for helpful and thorough comments. HIFI has been designed and built by a consortium of institutes and university departments from across Europe, Canada and the United States under the leadership of SRON Netherlands Institute for Space Research, Groningen, The Netherlands and with major contributions from Germany, France and the US. Consortium members are: Canada: CSA, U.Waterloo; France: CESR, LAB, LERMA, IRAM; Germany: KOSMA, MPIfR, MPS; Ireland, NUI Maynooth; Italy: ASI, IFSI-INAF, Osservatorio Astrofisico di Arcetri-INAF; Netherlands: SRON, TUD; Poland: CAMK, CBK; Spain: Observatorio Astronómico Nacional (IGN), Centro de Astrobiología (CSIC-INTA). Sweden: Chalmers University of Technology - MC2, RSS GARD; Onsala Space Observatory; Swedish National Space Board, Stockholm University - Stockholm Observatory; Switzerland: ETH Zurich, FHNW; USA: Caltech, JPL, NHSC. This work was supported by the Momentum grant of the MTA CSFK Lendület Disk Research Group and by JSPS KAKENHI Grant Number JP16F16770. This research has made use of NASA’s Astrophysics Data System.