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tokens: ACKNOWLEDGEMENTS We are grateful to Miklós Slíz (software engineer, Graphisoft, Budapest, Hungary) for the development of the computer simulation software. We thank Tatiana Salnikova for her valuable comments on an earlier version of our manuscript. Author contributions . Substantial contributions to conception and design: JSB, AB and GH. Performing experiments and data acquisition: JSB and AB. Data analysis and interpretation: JSB and GH. Drafting the article or revising it critically for important intellectual content: JSB and GH.