Holothyrida
# Holothyrida

The Holothyrida are a small order of mites in the superorder Parasitiformes. No fossils are known. With body lengths of more than 2 mm (3⁄32 in) they are relatively large mites, with a heavily sclerotized body. It is divided into three families, Allothyridae, Holothyridae, and Neothyridae. In a 1998 experimental study, members of the family Allothyridae were found to ignore living animals but readily fed on the body fluids of dead arthropods, making them scavengers.

The order has a distribution largely confined to former Gondwanan landmasses. They are the sister group to Ixodida (ticks).

## Systematics

### Allothyridae

Allothyridae van der Hammen, 1972 — Australia, New Zealand

* Allothyrus van der Hammen, 1961

   * Allothyrus australasiae (Womersley, 1935)

   * Allothyrus constrictus (Domrow, 1955)
* Australothyrus van der Hammen, 1983

   * Australothyrus ocellatus van der Hammen, 1983

### Holothyridae

Holothyridae Thorell, 1882 Sri Lanka, Indian Ocean islands, New Guinea, New Caledonia

* Sternothyrus Lehtinen, 1995
   * Sternothyrus braueri (Thon, 1905) — Seychelles
* Lindothyrus Lehtinen, 1995
   * Lindothyrus elongatus Lehtinen, 1995 — Lord Howe Island
   * Lindothyrus rubellus Lehtinen, 1995 — New Caledonia
* Indothyrus Lehtinen, 1995
   * Indothyrus greeni Lehtinen, 1995 — Sri Lanka
* Haplothyrus Lehtinen, 1995
   * Haplothyrus expolitissimus (Berlese, 1924) — New Caledonia
   * Haplothyrus hyatti Lehtinen, 1995 — unknown locality
* Holothyrus Gervais, 1842 — Mauritius
   * Holothyrus coccinella Gervais, 1842
   * Holothyrus legendrei Hammen, 1983
* Hammenius Lehtinen, 1981
   * Hammenius armatus (Canestrini, 1897) — Tamara Island (Aitape): New Guinea
   * Hammenius berlesei (Lehtinen, 1995) — New Guinea
   * Hammenius braueri (Thon, 1906)
   * Hammenius fujuge Lehtinen, 1981 — New Guinea (Central District, Oro Province)
   * Hammenius grandjeani (Hammen, 1961) — Mount Bosavi: New Guinea
   * Hammenius holthuisi van der Hammen, 1983
   * Hammenius ingii Lehtinen, 1981 — New Guinea
   * Hammenius insularis Lehtinen, 1995 — Louisiade Archipelago: New Guinea
   * Hammenius longipes (Thorell, 1882) — Fly River, New Guinea (?)
   * Hammenius mendi (Lehtinen, 1995) — Strickland River: New Guinea
   * Hammenius montanus Hammen, 1983 — Irian Jaya
   * Hammenius niger (Thon, 1906)

### Neothyridae

Neothyridae Lehtinen, 1981 Northern South America and the Caribbean

* Diplothyrus Lehtinen, 1999 Brazil, French Guiana
   * Diplothyrus schubarti Lehtinen, 1999
   * Diplothyrus lecorrei Klompen 2010
   * Diplothyrus lehtineni Vázquez & de Araújo & Feres 2016
* Neothyrus Lehtinen, 1981 Peru
   * Neothyrus ana Lehtinen, 1981
* Caribothyrus Kontschán & Mahunka 2004 Dominican Republic
   * Caribothyrus barbatus Kontschán & Mahunka 2004

## Footnotes

* ↑ Walter, David Evans; Proctor, Heather C. (1998). "Feeding behaviour and phylogeny: observations on early derivative Acari". Experimental and Applied Acarology. 22 (1): 39–50. doi:10.1023/A:1006033407957.
* ↑ Dobson, Susan J.; Barker, Stephen C. (1999). "Phylogeny of the Hard Ticks (Ixodidae) Inferred from 18S rRNA Indicates That the GenusAponommaIs Paraphyletic". Molecular Phylogenetics and Evolution. 11 (2): 288–295. doi:10.1006/mpev.1998.0565.
