Columella (genus)
# Columella (genus)

Columella is a genus of very small, air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Truncatellinidae, the whorl snails and allies.

## Taxonomy

Columella is commonly classified within the Vertiginidae. However, as shown by Nekola & Coles (2016), the genus is not that closely related to the Vertiginidae, but may rather be affiliated with the Chondrinidae or form a pupilloid family of its own. A systematic revision is pending.

## Species

Species within the genus Columella include:

* Columella acicularis Almuhambetova, 1979
* Columella aspera Waldén, 1966
* Columella columella (G. v. Martens, 1830)
* Columella edentula (Draparnaud, 1805) - toothless column
* †Columella hartmutnordsiecki Schlickum & Geissert, 1980
* Columella hasta (Hanna, 1911)
* Columella intermedia Schileyko & Almuhambetova, 1984
* Columella microspora (R.T. Lowe, 1852)
* Columella nymphaepratensis Hlaváč & Pokryszko, 2009
* Columella polvonense (Pilsbry, 1894)
* Columella simplex (Gould, 1840)
* Columella talgarica Schileyko & Rymzhanov, 2010
* †Columella wyciski Schlickum & Strauch, 1972

### Species brought into synonymy

* Columella inornata (Michaud, 1831): synonym of Columella edentula (Draparnaud, 1805) (junior synonym)
* Columella ninagongonis Pilsbry, 1935: synonym of Truncatellina ninagongonis (Pilsbry, 1935) (original combination)
* Columella tridentata A. B. Leonard, 1946: synonym of Gastrocopta ruidosensis (Cockerell, 1899)
