Orthogonius
# Orthogonius

Orthogonius is a genus of beetles in the family Carabidae, containing the following species:

* Orthogonius aberlenci Tian & Deuve, 2016 (Laos)
* Orthogonius acrogonus (Wiedemann, 1819) (Malaysia, Indonesia, and Borneo)
* Orthogonius acutangulus Chaudoir, 1878 (Sri Lanka)
* Orthogonius adoriae Tian & Deuve, 2006 (India)
* Orthogonius aemulus Péringuey, 1896 (Zimbabwe, Botswana, and Namibia)
* Orthogonius alternans (Wiedemann, 1823) (Indomalaya)
* Orthogonius alutaceus Quedenfeldt, 1883 (Africa)
* Orthogonius andrewesi Emden, 1928 (Philippines)
* Orthogonius andrewesianus Tian & Deuve, 2006 (Malaysia, Singapore, and Indonesia)
* Orthogonius angkor Tian & Deuve, 2006 (Cambodia)
* Orthogonius angkorensis Tian & Deuve, 2008 (Cambodia)
* Orthogonius angustus Chaudoir, 1872 (Thailand, Vietnam, and Malaysia)
* Orthogonius annamensis Jedlicka, 1965 (Vietnam)
* Orthogonius annamicola Tian & Deuve, 2006 (Vietnam)
* Orthogonius annemariae Tian & Deuve, 2006 (India)
* Orthogonius assamensis Tian & Deuve, 2006 (India)
* Orthogonius assamicola Tian & Deuve, 2006 (India)
* Orthogonius assmuthi Wasmann, 1920 (India)
* Orthogonius baconi Chaudoir, 1872 (India and Myanmar)
* Orthogonius baconioides Tian & Sabu, 2009 (India)
* Orthogonius balthasari Jedlicka, 1935 (Philippines)
* Orthogonius batesi Tian & Deuve, 2003 (Sri Lanka)
* Orthogonius batesianus Tian & Deuve, 2007
* Orthogonius batesioides Tian & Deuve, 2006 (India)
* Orthogonius baumi Jedlicka, 1955 (Malaysia)
* Orthogonius becqueti Burgeon, 1937 (Democratic Republic of the Congo and Burundi)
* Orthogonius bellus Tian & Deuve, 2016 (Vietnam)
* Orthogonius bicolor Jedlicka, 1965 (Vietnam)
* Orthogonius borneoensis Tian & Deuve, 2006 (Indonesia and Borneo)
* Orthogonius brancuccii Tian & Deuve, 2006 (Thailand)
* Orthogonius brevilabris Kolbe, 1889 (Democratic Republic of the Congo and Angola)
* Orthogonius brevithorax Dejean, 1825 (Africa)
* Orthogonius bruggei Tian & Deuve, 2006 (Malaysia)
* Orthogonius buquetii Chaudoir, 1850 (Sierra Leone and Ivory Coast)
* Orthogonius burckhardti Tian & Deuve, 2006
* Orthogonius burmanensis Tian & Deuve, 2006 (Myanmar)
* Orthogonius caffer Boheman, 1848 (Zimbabwe and South Africa)
* Orthogonius cambodgensis Tian & Deuve, 2006 (Cambodia)
* Orthogonius canaraensis Tian & Deuve, 2006 (India)
* Orthogonius canaraicola Tian & Deuve, 2006 (India)
* Orthogonius capucinus Boheman, 1848 (Zimbabwe and South Africa)
* Orthogonius carbonicolor Tian & Deuve, 2006 (Myanmar)
* Orthogonius carinatus Tian & Deuve, 2013 (Laos)
* Orthogonius celebesicus Tian & Deuve, 2006 (Indonesia)
* Orthogonius ceylanicus Tian & Deuve, 2016 (Sri Lanka)
* Orthogonius chaudoiri Tian & Deuve, 2001 (Vietnam)
* Orthogonius cheni Tian & Deuve, 2001 (China)
* Orthogonius chiangensis Tian & Deuve, 2006 (Thailand)
* Orthogonius clarkii Murray, 1858 (Africa)
* Orthogonius confusus Tian & Deuve, 2006 (Malaysia)
* Orthogonius constanti Tian & Deuve, 2006 (Cambodia)
* Orthogonius constrictus Tian; Deuve & Felix, 2012 (Thailand)
* Orthogonius coomani Tian & Deuve, 2006 (Vietnam)
* Orthogonius coomanioides Tian; Deuve & Felix, 2012 (Thailand)
* Orthogonius coracinus Kolbe, 1895 (Uganda, Rwanda, Burundi, and Tanzania)
* Orthogonius crassicrus Chaudoir, 1872 (Indonesia and Borneo)
* Orthogonius cruralis Putzeys in Chaudoir, 1872
* Orthogonius curvatus Tian & Deuve, 2006 (India)
* Orthogonius cyclothorax Tian & Deuve, 2007 (Singapore)
* Orthogonius davidi Chaudoir, 1878 (China and Taiwan)
* Orthogonius deletus Schmidt-Goebel, 1846 (China, India, Southeast Asia)
* Orthogonius deuvei Tian & Kirschenhofer, 2013 (Vietnam)
* Orthogonius dispar Bates, 1892 (Myanmar)
* Orthogonius dohertyi Tian & Deuve, 2006 (Malaysia)
* Orthogonius dongnanya Tian & Deuve, 2006 (Singapore, Indonesia, and Borneo)
* Orthogonius doriae Putzeys in Chaudoir, 1872 (Indonesia and Borneo)
* Orthogonius drescheri Liebke, 1937 (Indonesia)
* Orthogonius drumonti Tian & Deuve, 2006 (Indonesia)
* Orthogonius duboisi Tian & Deuve, 2006 (China)
* Orthogonius duplicatus (Wiedemann, 1819) (Indomalaya)
* Orthogonius dupuisi Basilewsky, 1948 (Zambia)
* Orthogonius dupuisianus Tian & Deuve, 2010 (Indonesia)
* Orthogonius dureli Tian & Deuve, 2005 (Bhutan and India)
* Orthogonius edentatus Tian & Deuve, 2006 (Malaysia, Indonesia, and Borneo)
* Orthogonius elisabethanus Burgeon, 1937 (Democratic Republic of the Congo)
* Orthogonius emarginatus Tian & Deuve, 2007 (Singapore)
* Orthogonius equimarginalis Tian & Deuve, 2006 (Indonesia)
* Orthogonius euthyphallus Tian & Deuve, 2013 (Laos and Vietnam)
* Orthogonius fairmairei Tian & Deuve, 2006 (Singapore)
* Orthogonius feai Tian & Deuve, 2007
* Orthogonius femoralis Chaudoir, 1848 (Sri Lanka and India)
* Orthogonius flavipes Deuve, 2004 (Indonesia)
* Orthogonius flavus Jedlicka, 1964 (India)
* Orthogonius foveiclypeus Tian & Deuve, 2004 (Indonesia)
* Orthogonius fryi Tian & Deuve, 2006 (Myanmar)
* Orthogonius fugax Chaudoir, 1872 (Sri Lanka)
* Orthogonius gracililamella Tian; Deuve & Felix, 2012 (Thailand)
* Orthogonius gracilipes Tian & Deuve, 2006 (India)
* Orthogonius gracilis Tian & Deuve, 2006 (India)
* Orthogonius grootaerti Tian & Deuve, 2006 (Thailand)
* Orthogonius hageni Oberthür, 1883 (Indonesia)
* Orthogonius himalaya Tian & Deuve, 2006 (Bhutan and India)
* Orthogonius himalayicus Tian & Deuve, 2005 (Bhutan and India)
* Orthogonius hirtulus Tian & Deuve, 2006 (Indonesia)
* Orthogonius hirtus Chaudoir, 1872 (Malaysia and Indonesia)
* Orthogonius hopei Gray, 1832 (Indomalaya)
* Orthogonius huananoides Tian & Deuve, 2006 (Thailand and Vietnam)
* Orthogonius huananus Tian & Deuve, 2001 (China)
* Orthogonius hypocrita Chaudoir, 1872 (Indonesia and Philippines)
* Orthogonius hypocritoides Jedlicka, 1935 (Philippines)
* Orthogonius impunctipennis Quedenfeldt, 1883 (Democratic Republic of the Congo and Angola)
* Orthogonius indicus Tian & Deuve, 2006 (India)
* Orthogonius indophilus Tian & Deuve, 2006 (India)
* Orthogonius inexpectatus Tian & Deuve, 2006 (India)
* Orthogonius inops Chaudoir, 1872 (Indonesia)
* Orthogonius insularis Chaudoir, 1872 (Malaysia, Indonesia, and Borneo)
* Orthogonius intermedius Chaudoir, 1872 (Indonesia)
* Orthogonius javanus Tian & Deuve, 2006 (Taiwan and Indonesia)
* Orthogonius jianfengling Tian & Deuve, 2001 (China)
* Orthogonius kalimantanensis Tian & Deuve, 2006 (Indonesia and Borneo)
* Orthogonius kapiriensis Burgeon, 1937 (Democratic Republic of the Congo)
* Orthogonius katangensis Burgeon, 1937 (Democratic Republic of the Congo)
* Orthogonius kickeli Kolbe, 1895 (Namibia)
* Orthogonius kirirom Tian & Deuve, 2008 (Thailand and Cambodia)
* Orthogonius klickai Jedlicka, 1935 (Philippines)
* Orthogonius kubani Tian & Deuve, 2006 (Thailand)
* Orthogonius kumatai Habu, 1979 (Nepal)
* Orthogonius lancangjiang Tian & Deuve, 2006 (Laos)
* Orthogonius lao Tian & Deuve, 2006 (Laos)
* Orthogonius latreillei Tian & Deuve, 2006 (India)
* Orthogonius latus Hope, 1842 (Africa)
* Orthogonius ledouxi Tian & Deuve, 2006 (Vietnam)
* Orthogonius legrandi Tian & Deuve, 2006 (Malaysia)
* Orthogonius lieftincki Andrewes, 1936 (Indonesia)
* Orthogonius limbourgi Tian & Deuve, 2016 (Vietnam)
* Orthogonius loeicus Tian & Deuve, 2006 (Thailand)
* Orthogonius loeiensis Tian & Deuve, 2006 (Thailand)
* Orthogonius longicornis Chaudoir, 1872 (Sri Lanka and Thailand)
* Orthogonius longilamella Tian & Deuve, 2008
* Orthogonius longipennis Hope, 1842
* Orthogonius longiphallus Tian & Deuve, 2005 (Bhutan)
* Orthogonius lucidus Bates, 1891 (India)
* Orthogonius luzonicus Chaudoir, 1872 (Philippines)
* Orthogonius macrophthalmus Tian & Deuve, 2013 (Vietnam)
* Orthogonius maindronianus Tian & Deuve, 2006 (India)
* Orthogonius makiling Tian & Deuve, 2006 (Philippines)
* Orthogonius malacca Tian & Deuve, 2006 (Malaysia)
* Orthogonius malaisei Andrewes, 1947 (Myanmar)
* Orthogonius malaya Tian & Deuve, 2006 (Malaysia)
* Orthogonius malaysiaensis Tian & Deuve, 2006 (Malaysia)
* Orthogonius medanensis Tian & Deuve, 2006 (Indonesia)
* Orthogonius meghalayaensis Tian & Deuve, 2016 (India)
* Orthogonius mekongensis Tian & Deuve, 2006 (Laos)
* Orthogonius melanipes Tian & Deuve, 2006 (India)
* Orthogonius mellyi (Chaudoir, 1850) (Bangladesh, India, and Myanmar)
* Orthogonius minimus Tian & Deuve, 2006 (India)
* Orthogonius mniszechi Chaudoir, 1872 (India, Malaysia, Indonesia, and Borneo)
* Orthogonius moestus Chaudoir, 1872
* Orthogonius monolophus Andrewes, 1931 (Indonesia and Borneo)
* Orthogonius montanus Tian & Deuve, 2006 (Myanmar)
* Orthogonius morvani Tian & Deuve, 2003 (Thailand)
* Orthogonius morvanianus Tian & Deuve, 2016 (Thailand)
* Orthogonius mouhoti Chaudoir, 1872 (Southeast Asia)
* Orthogonius myanmarensis Tian & Deuve, 2006 (Myanmar)
* Orthogonius nahaeo Tian & Deuve, 2006 (Thailand)
* Orthogonius nepalensis Habu, 1979 (Nepal)
* Orthogonius nepalicus Tian & Deuve, 2006 (Nepal)
* Orthogonius niger Jedlicka, 1935 (Philippines)
* Orthogonius nigripes Tian & Deuve, 2001 (China and Vietnam)
* Orthogonius nongfaensis Tian & Kirschenhofer, 2013 (Laos)
* Orthogonius nyassicus Kolbe, 1895
* Orthogonius obliquepes Tian & Deuve, 2006 (Philippines)
* Orthogonius opacus Schmidt-Goebel, 1846 (Indomalaya)
* Orthogonius orphnodes Andrewes, 1930 (Indonesia)
* Orthogonius ovatulus Tian & Deuve, 2003 (Sri Lanka)
* Orthogonius ovatus Chaudoir, 1878 (Indonesia)
* Orthogonius pachlatkoi Tian & Deuve, 2006 (Thailand)
* Orthogonius palangbangensis Tian & Deuve, 2006 (Indonesia)
* Orthogonius panghongae Tian & Deuve, 2006 (China)
* Orthogonius pangi Tian & Deuve, 2006 (Thailand and Vietnam)
* Orthogonius parallelus Chaudoir, 1872 (Sri Lanka)
* Orthogonius parastantoni Tian & Deuve, 2006 (Sri Lanka)
* Orthogonius parasuturalis Tian & Deuve, 2006 (Indonesia)
* Orthogonius parcepunctatus Kolbe, 1895 (Tanzania)
* Orthogonius paris Tian & Deuve, 2006 (Thailand)
* Orthogonius parvus Chaudoir, 1872 (India)
* Orthogonius pectinatus Tian & Deuve, 2016 (Thailand and Vietnam)
* Orthogonius perakensis Tian & Deuve, 2006 (Malaysia)
* Orthogonius perakicus Tian & Deuve, 2007 (Malaysia)
* Orthogonius perpuncticollis Burgeon, 1937 (Africa)
* Orthogonius petiolaris Tian & Deuve, 2006 (India)
* Orthogonius piceus Chaudoir, 1872 (Malaysia and Indonesia)
* Orthogonius picilabris (W.S.MacLeay, 1825) (Malaysia, Indonesia, and Borneo)
* Orthogonius picipennis Chaudoir, 1872 (Cambodia and Vietnam)
* Orthogonius pinguis Murray, 1858 (Nigeria)
* Orthogonius pinophilus Tian; Deuve & Felix, 2012 (Thailand)
* Orthogonius planiger (Walker, 1858) (Sri Lanka)
* Orthogonius plebius Tian & Deuve, 2006 (Sri Lanka)
* Orthogonius poggii Tian & Deuve, 2007
* Orthogonius politior Tian & Deuve, 2016 (Laos)
* Orthogonius politus Chaudoir, 1872 (Malaysia, Indonesia, and Borneo)
* Orthogonius pollinctor Basilewsky, 1948 (Democratic Republic of the Congo)
* Orthogonius pradieri Chaudoir, 1872
* Orthogonius pseudochaudoiri Tian; Deuve & Felix, 2012 (Thailand)
* Orthogonius pseudocheni Tian & Deuve, 2006 (Vietnam)
* Orthogonius pseudolongicornis Tian & Deuve, 2006 (Southeast Asia)
* Orthogonius pseudomouhoti Tian & Deuve, 2008 (Cambodia)
* Orthogonius punctatellus Tian & Deuve, 2006 (Indonesia)
* Orthogonius punctulatus Chaudoir, 1872 (India)
* Orthogonius punctum Tian & Deuve, 2016 (Thailand and Vietnam)
* Orthogonius putzeysi Tian & Deuve, 2006 (Sri Lanka)
* Orthogonius quadrisetosus Tian & Deuve, 2006 (Malaysia)
* Orthogonius rhodesianus Csiki, 1932
* Orthogonius rhodesiensis Lorenz, 1998 (Zambia)
* Orthogonius ribbei Tian & Deuve, 2003 (Indonesia)
* Orthogonius rogueti Tian & Deuve, 2016 (India)
* Orthogonius rotundatus Tian & Deuve, 2006
* Orthogonius rufiventris Bates, 1892 (Myanmar)
* Orthogonius rufotestaceus G.Müller, 1941 (Ethiopia)
* Orthogonius sabahensis Tian & Deuve, 2006 (Malaysia, Indonesia, and Borneo)
* Orthogonius sabahicus Tian & Deuve, 2016 (Malaysia, Indonesia, and Borneo)
* Orthogonius salvazai Tian & Deuve, 2006 (Laos)
* Orthogonius sarawakensis Tian & Deuve, 2006 (Malaysia, Indonesia, and Borneo)
* Orthogonius saundersi Andrewes, 1926 (Malaysia and Singapore)
* Orthogonius schaumi Chaudoir, 1872 (Sri Lanka and Indonesia)
* Orthogonius schaumioides Tian & Deuve, 2006 (India)
* Orthogonius schauteni Tian & Deuve, 2006 (Malaysia)
* Orthogonius schmidtgoebeli Chaudoir, 1872 (Myanmar)
* Orthogonius senegalensis Dejean, 1831 (Africa)
* Orthogonius setosopalpiger Tian; Deuve & Felix, 2012 (Thailand)
* Orthogonius siamensis Tian & Deuve, 2006 (Thailand)
* Orthogonius similaris Tian; Deuve & Felix, 2012 (Thailand)
* Orthogonius simplicatus Tian & Deuve, 2006 (Indonesia)
* Orthogonius singaporensis Tian & Deuve, 2006 (Singapore)
* Orthogonius sinuatiphallus Tian & Deuve, 2001 (China)
* Orthogonius smetsi Tian & Deuve, 2006 (Cambodia)
* Orthogonius solidicornis Tian & Deuve, 2003 (China)
* Orthogonius solidus Tian & Deuve, 2006 (India)
* Orthogonius spinosus Burgeon, 1937 (Africa)
* Orthogonius srilankaicus Tian & Deuve, 2003 (Sri Lanka)
* Orthogonius stantoni Tian & Deuve, 2006 (India)
* Orthogonius sterbai Jedlicka, 1935 (Philippines)
* Orthogonius stygius Andrewes, 1930 (Malaysia and Indonesia)
* Orthogonius sulawesicus Tian & Deuve, 2003 (Indonesia)
* Orthogonius sulcatoides Tian & Deuve, 2006 (India, Myanmar, and Malaysia)
* Orthogonius sulcatus Schmidt-Goebel, 1846 (Myanmar and Cambodia)
* Orthogonius sumatraicus Tian & Deuve, 2006 (Indonesia)
* Orthogonius suturalis Chaudoir, 1872 (Malaysia, Indonesia, and Borneo)
* Orthogonius taghavianae Tian; Deuve & Felix, 2012 (Thailand)
* Orthogonius taiwanicus Tian & Deuve, 2006 (Taiwan)
* Orthogonius tangsen Tian & Deuve, 2006 (India)
* Orthogonius tenasserimensis Tian & Deuve, 2006 (Myanmar)
* Orthogonius termiticola Wasmann, 1902 (Malaysia)
* Orthogonius thaicus Tian & Deuve, 2003 (Thailand)
* Orthogonius thaiensis Tian & Deuve, 2006 (Thailand)
* Orthogonius thailandensis Tian & Deuve, 2006 (Thailand)
* Orthogonius thoracicus Gestro, 1875
* Orthogonius tikekee Tian & Deuve, 2006 (Myanmar)
* Orthogonius tonkinensis Tian & Deuve, 2006 (Vietnam)
* Orthogonius tonkinicus Tian & Deuve, 2006 (Vietnam)
* Orthogonius travancore Tian & Deuve, 2006 (India)
* Orthogonius tricarinatus Burgeon, 1937 (Africa)
* Orthogonius uncipennis Tian & Deuve, 2006 (Nepal)
* Orthogonius vari Tian; Deuve & Felix, 2012 (Thailand and Cambodia)
* Orthogonius variabilis Tian; Deuve & Felix, 2012 (China and Thailand)
* Orthogonius virgulatus Andrewes, 1931 (Indonesia and Borneo)
* Orthogonius wallardiensis Tian & Deuve, 2006 (India)
* Orthogonius wrasei Tian & Deuve, 2016 (Myanmar)
* Orthogonius xanthomerus L.Redtenbacher, 1868 (China)
* Orthogonius yoga Tian & Deuve, 2006 (India)
* Orthogonius yunnanensis Tian & Deuve, 2001 (China)
