Arthropods of the Cambrian Period
# Arthropods of the Cambrian Period

This list contains extinct arthropod genera from the Cambrian Period of the Paleozoic Era.

## Artiopoda

| Genus | Year of Discovery | Location of Fossils |
| - | - | - |
| Acanthomeridion | 1989 | Chengjiang Biota |

## Dinocaridida

| Genus | Year of Discovery | Location of Fossils |
| - | - | - |
| Anomalocaris | 1892 | Emu Bay Shale Maotianshan Shales Burgess Shale |
| Cucumericrus | 1995 | Chengjiang Biota |
| Paranomalocaris | 2013 | Wulongqing Formation |
| Cambroraster | 2019 | Burgess Shale Chengjiang Biota |
| Hurdia | 1992 | Burgess Shale |
| Peytoia | 1911 | Burgess Shale Zawiszyn Formation |
| Ursulinacaris | 2019 | Carrara Formation |
| Titanokorys | 2018 | Burgess Shale |
| Pahvantia | 1981 | Wheeler Shale Marjum Formation |
| Amplectobelua | 1995 | Chengjiang Burgess Shale |
| Houcaris | 2021 | |
| Lyrarapax | 2014 | |
| Caryosyntrips | 2010 | Burgess Shale Wheeler Shale Marjum Formation Valdemiedes Formation |
| Tamisiocaris | 2010 | Buen Formation Sirius Passet |
| Opabinia | 1912 | Burgess Shale |
| Utaurora | 2022 | Wheeler Formation |
| Kerygmachela | 1993 | Buen Formation Sirius Passet |
| Laminacaris | 2018 | |
| Pambdelurion | 1997 | Sirius Passet |

## Malacostraca

| Genus | Year of Discovery | Location of Fossils |
| - | - | - |
| Hymenocaris | 1853 | Burgess Shale |
| Odaraia | 1912 | Burgess Shale Greater Phyllopod Bed |

## Mandibulata

| Genus | Year of Discovery | Location of Fossils |
| - | - | - |
| Branchiocaris | 1929 | Wheeler Shale Burgess Shale |
| Tokummia | 2017 | Burgess Shale |
| Canadaspis | 1978 | Burgess Shale Pioche Shale Chengjiang Biota |
| Waptia | 1912 | Burgess Shale |
| Pectocaris | 1999 | Maotianshan Shales Burgess Shale |
| Clypecaris | 1999 | Chengjiang Biota |
| Pauloterminus | 2002 | Sirius Passet |
| Perspicaris | 1977 | Burgess Shale |
| Nereocaris | 2012 | Burgess Shale |
| Plenocaris | 1912 | Burgess Shale Chengjiang Biota |
| Ercaicunia | 1999 | Maotianshan Shales |
| Fibulacaris | 2019 | Burgess Shale |
| Synophalos | 2009 | Maotianshan Shales |

## Marrellomorpha

| Genus | Year of Discovery | Location of Fossils |
| - | - | - |
| Marrella | 1912 | Burgess Shale Kaili Formation Balang Formation |
| Skania | 1931 | Maotianshan Shales Burgess Shale |
| Primicaris | 2003 | Chengjiang Biota |

## Maxillopoda

| Genus | Year of Discovery | Location of Fossils |
| - | - | - |
| Heymonsicambria | 1994 | Orsten Fauna |
| Skara | 1983 | Orsten Fauna |
| Goticaris | 1990 | Orsten Fauna |

## Megacheira

| Genus | Year of Discovery | Location of Fossils |
| - | - | - |
| Actaeus | 1970 | Burgess Shale |
| Alalcomenaeus | 1970 | Burgess Shale Chengjiang Biota Wheeler Shale |
| Leanchoilia | 1912 | Burgess Shale Chengjiang Biota Spence Shale |
| Fortiforceps | 1997 | Maotianshan Shales |
| Yawunik | 2015 | Burgess Shale |
| Parapeytoia | 1995 | Maotianshan Shales |
| Jianfengia | 1987 | Maotianshan Shales |
| Haikoucaris | 2004 | Chengjiang Biota |
| Kanoshoia | 2020 | Wheeler Formation |
| Yohoia | 1912 | Burgess Shale |

## Ostracoda

| Genus | Year of Discovery | Location of Fossils |
| - | - | - |
| Carnarvonia | 1912 | Burgess Shale |

## Trilobita

| Genus | Year of Discovery | Location of Fossils |
| - | - | - |
| Bathyuriscus | 1873 | Burgess Shale |
| Olenoides | 1877 | Burgess Shale Marjum Formation |
| Hanburia | 1916 | Burgess Shale Stephen Formation |
| Xiuqiella | 1974 | |
| Acontheus | 1851 | |
| Hemirhodon | 1937 | |
| Thoracocare | 1974 | Spence Shale Formation |
| Abakania | 1973 | Sanashtykgol Formation |
| Zacanthoides | 1888 | Burgess Shale |
| Changaspis | 1961 | |
| Ogygopsis | 1889 | Burgess Shale |
| Oryctocephalus | 1886 | Burgess Shale |
| Chancia | 1924 | Burgess Shale |
| Ehmaniella | 1951 | Burgess Shale |
| Elrathia | 1924 | Wheeler Formation Burgess Shale |
| Ellipsocephalus | 1883 | Słowiec Sandstone Formation |
| Phantaspis | 2020 | Mantou Formation |
| Aojia | 1935 | |
| Bailiaspis | 1936 | Whitesands Bay Formation Jbel Lmgaysmat Formation Manuels River Formation Maentwrog Formations |
| Solenoparia | 1935 | |
| Westergaardites | 1937 | |
| Yunnanocephalus | 1936 | Chengjiang Biota |
| Asaphiscus | 1873 | |
| Olenus | 1827 | |
| Quitacetra | 1967 | |
| Ejinaspis | 1986 | |
| Asthenopsis | 1939 | |
| Agraulos | 1847 | Menevia Formation Jince Formation Manuels River Formation |
| Coosella | 1936 | |
| Norwoodia | 1916 | |
| Tricrepicephalus | 1935 | Chefu Formation Deadwood Formation Little Elk Creek |
| Meniscopsia | 2011 | |
| Acrocephalella | 1963 | |
| Cambroproteus | 2015 | |
| Cedaria | 1924 | Weeks Formation |
| Conocoryphe | 1847 | |
| Galeaspis | 1983 | |
| Genevievella | 1936 | Mungerbar Formation |
| Meteoraspis | 1890 | Limestone Formation |
| Validaspis | 1991 | |
| Palaeolenus | 1912 | |
| Ptychoparia | 1847 | |
| Kootenia | 1939 | Burgess Shale Emu Bay Shale |
| Misszhouia | 1985 | Chengjiang Biota |
| Naraoia | 1912 | Burgess Shale Stephen Formation Chengjiang biota Formation of Southern Ontario Balang Formation Emu Bay Shale |
| Estaingia | 1964 | |
| Bathynotus | 1860 | |
| Bajanaspis | 1978 | Edreisk Formation |
| Kleptothule | 1995 | Sirius Passet |
| Acadoparadoxides | 1957 | |
| Paradoxides | 1822 | Duda Formation |
| Eoredlichia | 1950 | Chengjiang Biota |
| Profallotaspis | 2002 | |
| Lochmanolenellus | 1952 | Poleta Formation |
| Olenellus | 1861 | Pioche Shale Carrara Formation Kinzers Formation |
| Archaeaspis | 1965 | Campito Formation |
| Parapaokannia | 1980 | |
| Plutonides | 1985 | Purley Shale Formation |
| Redlichia | 1902 | |
| Repinaella | 1965 | Pestrotsvet Formation |
| Ushbaspis | 1965 | Shabakty Formation |
| Balcoracania | 1970 | Balcoracana Creek |
| Emuella | 1970 | Emu Bay |
| Megapharanaspis | 2006 | |
| Wutingaspis | 1944 | |
| Mesolenellus | 1993 | Sofiekammen Formation |
| Olenelloides | 1894 | |
| Biceratops | 1971 | Pioche Shale |
| Bradyfallotaspis | 1972 | |
| Eopeachella | 2009 | Delamar Mountains |
| Laudonia | 1956 | |
| Mummaspis | 1992 | Mural Formation |
| Nephrolenellus | 1993 | Pyramid Shale Member Carrara Formation |
| Nevadella | 1936 | Poleta Formation Sekwi Formation |
| Paranephrolenellus | 2007 | |
| Wanneria | 1910 | Kinzers Formation |
| Bristolia | 1956 | Carrara Formation |
| Acadoparadoxides | 1957 | |
| Emigrantia | 1910 | |
| Bolbolenellus | 1993 | |
| Buenellus | 1988 | Sirius Passet |
| Callavia | 1890 | Brigus Formation |
| Chengjiangaspis | 1980 | Maotianshan Shales |
| Cirquella | 1993 | |
| Daguinaspis | 1950 | Amouslek Formation |
| Fallotaspis | 1953 | |
| Holmia | 1890 | Sparagmite Formation |
| Hydrocephalus | 1846 | |
| Fritzolenellus | 1998 | |
| Mesonacis | 1859 | Forteau Formation |
| Xystridura | 1936 | |
| Peachella | 1910 | |
| Richterops | 1954 | |
| Archaeagnostus | 1939 | |
| Agnostus | 1822 | Marjum Biota Linyi Lagerstätte Orsten Fauna |
| Itagnostus | 1979 | |
| Peronopsis | 1847 | |
| Acmarhachis | 1938 | |
| Acadagnostus | 1939 | |
| Agnostotes | 1963 | |
| Lejopyge | 1828 | |
| Glyptagnostus | 1936 | |
| Egyngolia | 1980 | |
| Jinghediscus | 1985 | |
| Pagetia | 1916 | Burgess Shale |
| Acimetopus | 1966 | |
| Pentagnostus | 1940 | |
| Calodiscus | 1935 | Schodack Formation |
| Mallagnostus | 1935 | |
| Runcinodiscus | 1976 | |
| Serrodiscus | 1941 | Brigus Formation |
| Weymouthia | 1913 | |
| Analox | 1966 | |
| Litometopus | 1966 | |
| Meniscuchus | 1975 | |
| Ninadiscus | 1980 | Egyngolskaya Suite |
| Oodiscus | 1966 | |
| Semadiscus | 1978 | |
| Stigmadiscus | 1966 | |
| Parapagetia | 1964 | |
| Tannudiscus | 1959 | Purley Shale Formation |
| Chelediscus | 1966 | |
| Bathydiscus | 1966 | |
| Cobboldites | 1943 | |
| Acidiscus | 1966 | |
| Buenaspis | 1999 | Sirius Passet |
| Condylopyge | 1847 | |
| Delgadella | 1908 | Tyuser Formation |
| Dicerodiscus | 1954 | |
| Pleuroctenium | 1847 | |
| Ptychagnostus | 1909 | |
| Bolboparia | 1966 | |
| Tchernyshevioides | 1975 | |
| Toragnostus | 1988 | |
| Tsunyidiscus | 1966 | |
| Birmanites | 1934 | |
| Kendallina | 1953 | Bison Creek Formation |
| Dikelocephalus | 1852 | |
| Orygmaspis | 1938 | Morgan Creek Member |
| Cheirurus | 1845 | |
| Cindarella | 1996 | Chengjiang Biota |
| Helmetia | 1912 | Burgess Shale Jince Formation |
| Tegopelte | 1975 | Burgess Shale Kicking Horse Shale |
| Retifacies | 1989 | Maotianshan Shales |
| Schmalenseeia | 1903 | |
| Burlingia | 1908 | |
| Entomaspis | 1931 | |
| Neodrepanura | 2006 | Maotianshan Shales |

## Incertae sedis

| Genus | Year of Discovery | Location of Fossils |
| - | - | - |
| Kiisortoqia | 2010 | Sirius Passet |
| Vladicaris | 1995 | |
| Wujicaris | 2010 | Orsten Fauna |
| Protocaris | 1884 | Parker Formation |
| Burgessia | 1912 | Burgess Shale |
| Liangshanella | 1997 | Chengjiang Biota Burgess Shale |
| Kunmingella | 1912 | Maotianshan Shales |
| Comptaluta | 1968 | Maotianshan Shales |
| Jiucunella | 1991 | Maotianshan Shales |
| Kunyangella | 1965 | Maotianshan Shales |
| Cyclopites | 1951 | |
| Mongolitubulus | 1997 | |
| Habelia | 1912 | Burgess Shale |
| Khankaspis | 1969 | Snegurovka Formation |
| Isoxys | 1890 | Burgess Shale Chengjiang Biota Sirius Passet |
| Emeraldella | 1912 | Burgess Shale Wheeler Shale |
| Molaria | 1912 | Burgess Shale Sirius Passet |
| Sidneyia | 1911 | Burgess Shale Maotianshan Shales |
| Bailongia | 2021 | Maotianshan Shales |
| Luohuilinella | 2012 | Chengjiang Biota |
| Panlongia | 2006 | Guanshan Fauna |
| Pygmaclypeatus | 2000 | Chengjiang Biota |
| Sinoburius | 1991 | Chengjiang Biota |
| Retifacies | 1989 | Chengjiang Biota |
| Strabops | 1901 | |
| Paleomerus | 1956 | |
| Parapaleomerus | 1999 | Chengjiang Biota |
| Sanctacaris | 1988 | Burgess Shale |
| Mollisonia | | Chengjiang Biota Burgess Shale Langston Formation Wheeler Shale |
| Thelxiope | 1975 | Burgess Shale Wheeler Shale Fezouata Formation Linyi Lagerstätte |
| Glypharthrus | 1939 | McKay Group Clay Creek |
| Urokodia | 1989 | Maotianshan Shales |
| Fuxianhuia | 1987 | Chengjiang Biota |
| Alacaris | 2018 | |
| Chengjiangocaris | 1991 | |
| Guangweicaris | 2007 | |
| Tuzoia | 1912 | Burgess Shale Emu Bay Shale Rockslide Formation Spence Shale Wheeler Formation Marjum Formation Kinzers Formation |
| Klausmuelleria | 2003 | Orsten Fauna |
| Kylinxia | 2020 | Maotianshan Shales |
| Aaveqaspis | 2009 | Sirius Passet |
| Worthenella | 1911 | Burgess Shale |
| Kodymirus | 1965 | Paseky Shale |
| Mosineia | 2010 | Blackberry Hill |
| Cambropachycope | 1990 | Orsten Fauna |
| Bushizheia | 2020 | Chiungchussu |
| Combinivalvula | 1987 | Chengjiang Biota |
| Diplopyge | 1999 | Chengjiang Biota |
| Dongshanocaris | 1999 | Chengjiang Biota |
| Ercaia | 2001 | Chengjiang Biota |
| Jianshania | 1999 | Chengjiang Biota |
| Pseudoiulia | 1998 | Chengjiang Biota |
| Priscansermarinus | 1981 | Burgess Shale |
| Kootenichela | 2013 | |
| Forfexicaris | 1999 | Maotianshan Shales |
| Kockurus | 1995 | |
| Occacaris | 1999 | Maotianshan Shales |
| Sarotrocercus | 1981 | Burgess Shale |
