Helius (fly)
# Helius (fly)

Helius is a genus of crane fly in the family Limoniidae.

## Distribution

Worldwide, but the Australasian and Oriental regions are the most species rich.

## Species

* Subgenus Eurhamphidia Alexander, 1915

   * H. abnormalis (Brunetti, 1918)

   * H. ata Alexander, 1931

   * H. atroapicalis Alexander, 1978

   * H. auranticolor Alexander, 1936

   * H. connectus Edwards, 1928

   * H. diacanthus Alexander, 1931

   * H. fuscofemoratus Alexander, 1931

   * H. glabristylatus Alexander, 1931

   * H. indivisus Alexander, 1931

   * H. inelegans (Alexander, 1928)

   * H. invenustipes (Alexander, 1930)

   * H. melanosoma Alexander, 1936

   * H. mimicans Edwards, 1933

   * H. mirus Edwards, 1926

   * H. monticola (Edwards, 1919)

   * H. mouensis Alexander, 1948

   * H. nigrofemoratus (Alexander, 1930)

   * H. niveitarsis (Skuse, 1890)

   * H. pallens Edwards, 1933

   * H. perelegans Alexander, 1930

   * H. perlongatus Alexander, 1978

   * H. scabiosus Alexander, 1932

   * H. vitiensis Alexander, 1956
* Subgenus Helius Lepeletier & Serville, 1828

   * H. acanthostyla Alexander, 1944

   * H. aciferus Alexander, 1937

   * H. aka Alexander, 1972

   * H. albitarsis (Osten Sacken, 1888)

   * H. albogeniculatus Alexander, 1936

   * H. amplus Edwards, 1933

   * H. anaemicus Alexander, 1932

   * H. anamalaiensis Alexander, 1967

   * H. angustalbus Alexander, 1953

   * H. aphrophilus Alexander, 1948

   * H. apicalis (Alexander, 1915)

   * H. apoensis Alexander, 1931

   * H. apophysalis Alexander, 1967

   * H. araucariae Alexander, 1945

   * H. arcuarius Alexander, 1929

   * H. argyrosternus Alexander, 1930

   * H. arunachalus Alexander, 1975

   * H. attenuatus Alexander, 1929

   * H. barbatus Edwards, 1921

   * H. bicolor Edwards, 1933

   * H. bifurcus Alexander, 1956

   * H. bitergatus Alexander, 1950

   * H. boops Alexander, 1942

   * H. brachyphallus Alexander, 1956

   * H. brevioricornis (Alexander, 1920)

   * H. brevisector Alexander, 1956

   * H. brunneus Byers, 1981

   * H. cacoxenus (Alexander, 1920)

   * H. calviensis Edwards, 1928

   * H. capensis (Alexander, 1917)

   * H. capniopterus Alexander, 1945

   * H. catreus Alexander, 1967

   * H. cavernicolus Alexander, 1961

   * H. chikurinensis Alexander, 1930

   * H. chintoo Theischinger, 1994

   * H. communis (Skuse, 1890)

   * H. comoreanus Alexander, 1959

   * H. compactus Alexander, 1967

   * H. copiosus Alexander, 1935

   * H. corniger Savchenko, 1983

   * H. costofimbriatus Alexander, 1930

   * H. costosetosus Alexander, 1932

   * H. creper Alexander, 1926

   * H. ctenonycha Alexander, 1938

   * H. dafla Alexander, 1972

   * H. darlingtonae Welch & Gelhaus, 1994

   * H. destitutus Alexander, 1971

   * H. devinctus Alexander, 1932

   * H. distinervis Alexander, 1940

   * H. dolichorhynchus Edwards, 1933

   * H. eremnophallus Alexander, 1978

   * H. euryphallus Alexander, 1960

   * H. fasciventris Edwards, 1926

   * H. ferruginosus (Brunetti, 1912)

   * H. flavidibasis Alexander, 1975

   * H. flavipes (Macquart, 1855)

   * H. flavitarsis (Alexander, 1920)

   * H. flavus (Walker, 1856)

   * H. fragosus Alexander, 1937

   * H. franckianus Alexander, 1940

   * H. fratellus (Brunetti, 1918)

   * H. fulvithorax (Skuse, 1890)

   * H. fumicosta Edwards, 1928

   * H. fuscoangustus Alexander, 1967

   * H. garcianus Alexander, 1972

   * H. gibbifer Savchenko, 1981

   * H. gorokanus Alexander, 1962

   * H. gracillimus Alexander, 1938

   * H. graphipterus Alexander, 1954

   * H. haemorrhoidalis Alexander, 1937

   * H. harrisi (Alexander, 1923)

   * H. hatschbachi Alexander, 1954

   * H. hispanicus Lackschewitz, 1928

   * H. hova Alexander, 1955

   * H. impensus Alexander, 1967

   * H. imperfectus (Alexander, 1920)

   * H. inconspicuus (Brunetti, 1912)

   * H. ineptus Alexander, 1938

   * H. infirmus Alexander, 1932

   * H. invariegatus Alexander, 1938

   * H. iris (Alexander, 1920)

   * H. kambangani (de Meijere, 1913)

   * H. kambanganoides Alexander, 1931

   * H. larotypa Alexander, 1926

   * H. lectus Alexander, 1936

   * H. leucoplaca Alexander, 1936

   * H. lienpingensis Alexander, 1945

   * H. liguliferus Alexander, 1978

   * H. liliputanus Alexander, 1929

   * H. lobuliferus Alexander, 1938

   * H. longinervis Edwards, 1928

   * H. longirostris (Meigen, 1818)

   * H. luachimoensis Alexander, 1963

   * H. luniger (Riedel, 1914)

   * H. mainensis (Alexander, 1916)

   * H. malagasicus Alexander, 1955

   * H. manueli Alexander, 1972

   * H. medleri Alexander, 1976

   * H. melanophallus Alexander, 1938

   * H. mesorhynchus Alexander, 1933

   * H. micracanthus Alexander, 1945

   * H. mirabilis (Alexander, 1914)

   * H. mirandus (Alexander, 1921)

   * H. morosus (Alexander, 1920)

   * H. multivolus Alexander, 1945

   * H. murreensis Alexander, 1960

   * H. myersiellus Alexander, 1935

   * H. nawaianus Alexander, 1929

   * H. neocaledonicus Alexander, 1945

   * H. nigricapella Alexander, 1938

   * H. nigriceps (Edwards, 1916)

   * H. nipponensis (Alexander, 1913)

   * H. numenius Alexander, 1967

   * H. obliteratus (Alexander, 1920)

   * H. obsoletus (Alexander, 1920)

   * H. oxystylus Alexander, 1967

   * H. pallidipes Alexander, 1926

   * H. pallidissimus Alexander, 1930

   * H. pallirostris Edwards, 1921

   * H. panamensis Alexander, 1934

   * H. paramorosus Alexander, 1949

   * H. parvidens Alexander, 1944

   * H. patens Edwards, 1933

   * H. pauperculus (Alexander, 1921)

   * H. pavoninus Alexander, 1938

   * H. perflavens Alexander, 1964

   * H. perpallidus Alexander, 1942

   * H. pervenustus Alexander, 1953

   * H. phasmatis Alexander, 1945

   * H. pictus Edwards, 1926

   * H. plebeius Alexander, 1946

   * H. pluto Alexander, 1932

   * H. polionotus Alexander, 1938

   * H. procerus Alexander, 1931

   * H. productellus Alexander, 1944

   * H. protumidus Alexander, 1978

   * H. quadrifidus Alexander, 1926

   * H. quadrivena Alexander, 1971

   * H. rectispinus Alexander, 1947

   * H. rectus Alexander, 1945

   * H. regius Alexander, 1944

   * H. rostratus Alexander, 1956

   * H. rubicundus (Alexander, 1922)

   * H. rufescens (Edwards, 1916)

   * H. rufithorax Alexander, 1928

   * H. sanguinolentus (Alexander, 1921)

   * H. schildi Alexander, 1945

   * H. selectus Alexander, 1936

   * H. serenus Alexander, 1967

   * H. setigerus Alexander, 1980

   * H. sigillatus Alexander, 1964

   * H. stenorhynchus Alexander, 1954

   * H. stolidus Alexander, 1948

   * H. subanaemicus Alexander, 1964

   * H. subarcuarius Alexander, 1934

   * H. subfasciatus Alexander, 1924

   * H. submorosus (Alexander, 1921)

   * H. subobsoletus (Alexander, 1921)

   * H. subpauperculus Alexander, 1975

   * H. tantalus Alexander, 1967

   * H. tanyrhinus Alexander, 1964

   * H. taos Alexander, 1967

   * H. tenuirostris Alexander, 1924

   * H. tenuistylus Alexander, 1929

   * H. tetracradus Alexander, 1967

   * H. tienmuanus Alexander, 1940

   * H. tonaleah Theischinger, 1994

   * H. trianguliferus Alexander, 1931

   * H. tshinta Theischinger, 1994

   * H. unicolor (Brunetti, 1912)

   * H. uniformis (Alexander, 1914)

   * H. venustus (Skuse, 1890)

   * H. verticillatus Alexander, 1967

   * H. yindi Theischinger, 1994
* Subgenus Idiohelius Alexander, 1948

   * H. melanolithus Alexander, 1953

   * H. mirificus Alexander, 1953

   * H. pentaneura Alexander, 1948
* Subgenus Mammuthonasus Theischinger, 1994

   * H. allunga Theischinger, 1994
* Subgenus Prohelius Alexander, 1975

   * H. dugaldi Alexander, 1945

   * H. edwardsianus Alexander, 1956

   * H. fulani Alexander, 1975
* Subgenus Rhamphidina Alexander, 1920

   * H. camerounensis (Alexander, 1920)
* Subgenus Rhamphidioides Alexander, 1920

   * H. alluaudi (Riedel, 1914)

   * H. venustissimus (Alexander, 1920)
* Subgenus Rhampholimnobia Alexander, 1915

   * H. bigeminatus Alexander, 1962

   * H. brevinasus Alexander, 1934

   * H. deentoo Theischinger, 1994

   * H. diffusus Alexander, 1941

   * H. fenestratus Alexander, 1973

   * H. gracilirostris Alexander, 1962

   * H. guttulinus Alexander, 1960

   * H. japenensis Alexander, 1948

   * H. matilei Hynes, 1993

   * H. mesolineatus Alexander, 1962

   * H. nimbus Alexander, 1941

   * H. papuanus Alexander, 1934

   * H. reticularis (Alexander, 1915)

   * H. simulator Alexander, 1962

   * H. spinteris Alexander, 1960

   * H. subreticulatus Alexander, 1962
* Subgenus Rhyncholimonia Alexander, 1964

   * H. dicroneurus Alexander, 1964
