Pterostichinae
# Pterostichinae

Pterostichinae Bonelli, 1810 è una sottofamiglia di coleotteri carabidi diffusa in tutto il mondo.

## Descrizione

Gli Pterostichinae sono coleotteri di medie o piccole dimensioni, forma ovale e colore scuro, spesso metallico. Mancano quasi completamente specie dotate di decorazioni a colori contrastanti. I caratteri che permettono di distinguerli dagli altri carabidi sono: le tibie anteriori con una incavatura all'apice, le elitre a curvatura regolare, le antenne pubescenti dal 4° articolo e gli occhi sormontati da due setole. Quest'ultima caratteristica permette di riconoscerli facilmente dagli Harpalinae, con cui possono venire confusi.

## Sistematica

La sottofamiglia è suddivisa in otto tribù che comprendono i seguenti generi:

* Tribù Abacetini Chaudoir, 1872
   * Abacetus Dejean, 1828
   * Abacidus LeConte, 1873
   * Aristopus Laferte-Senectere, 1853
   * Celioinkosa Straneo, 1951
   * Chlaeminus Motschulsky, 1865
   * Colpodichius Straneo, 1952
   * Cosmodiscus Sloane, 1907
   * Cyrtomoscelis Chaudoir, 1874
   * Distrigidius Jeannel, 1948
   * Ecnomolaus Bates, 1892
   * Haptoderidius Straneo, 1975
   * Haptoderodes Straneo, 1986
   * Holconotus Schmidt-Goebel, 1846
   * Inkosa Peringuey, 1926
   * Mateuellus Deuve, 1990
   * Metabacetus Bates, 1892
   * Metaxellus Straneo, 1960
   * Metaxys Chaudoir, 1857
   * Novillidius Straneo, 1941
   * Oodinkosa Straneo, 1939
   * Ophonichius Straneo, 1942
   * Pediomorphus Chaudoir, 1878
   * Pioprosopus Tschitscherine, 1899
   * Pollicobius Vinson, 1939
   * Prostalomus Basilewsky, 1950
   * Pseudabacetus Burgeon, 1935
   * Pterostillichus Straneo, 1949
   * Rhagadillius Straneo, 1951
   * Tiferonia Darlington, 1962
   * Trachelocyphoides Alluaud, 1934
   * Trachelocyphus Tschitscherine, 1900
* Tribù Chaetodactylini Tschitscherine, 1903
   * Chaetodactyla Tschitscherine, 1897
* Tribù Cnemalobini Germain, 1911
   * Cnemalobus Guerin-Meneville, 1838
* Tribù Cratocerini Lacordaire, 1854
   * Sottotribù Catapieseina Bates, 1882
      * Catapiesis Solier, 1835
      * Homalomorpha Brulle, 1835
   * Sottotribù Cratocerina sensu stricto
      * Brachidius Chaudoir, 1852
      * Cratocerus Dejean, 1829
      * Oxyglychus Straneo, 1938
   * Sottotribù Drimostomatina Chaudoir, 1872
      * Abacaelostus Straneo, 1952
      * Andrewesinulus Straneo, 1938
      * Apsidocnemus Alluaud, 1936
      * Barylaus Liebherr, 1985
      * Caecocaelus Straneo, 1949
      * Caelostomus W.S. MacLeay, 1825
      * Camptogenys Tschitscherine, 1899
      * Capabatus Csiki, 1930
      * Crenulostrigus Straneo, 1942
      * Cyrtolaus Bates, 1882
      * Dactyleurys Tschitscherine, 1899
      * Dactylinius Straneo, 1941
      * Diachipteryx Alluaud, 1926
      * Diceromerus Chaudoir, 1872
      * Dromistomus Jeannel, 1948
      * Drymonaxus Straneo, 1941
      * Eurystomis Chaudoir, 1878
      * Feostoma Straneo, 1941
      * Hemitelestus Alluaud, 1895
      * Hoplizomenus Chaudoir, 1872
      * Leleuporites Straneo, 1960
      * Madapelmus Dajoz, 1985
      * Monodryxus Straneo, 1942
      * Pachycaecus Straneo, 1971
      * Pachyroxochus Straneo, 1942
      * Platyxythrius Straneo, 1942
      * Stegazopteryx Will, 2004
      * Stomonaxellus Tschitscherine, 1901
      * Strigomerodes Straneo, 1939
      * Strigomerus Chaudoir, 1872
      * Trichillinus Straneo, 1938
* Tribù Microcheilini Jeannel, 1948
   * Microcheila Brulle, 1834
* Tribù Morionini Brulle, 1835
   * Buderes Murray, 1857
   * Hyperectenus Alluaud, 1935
   * Hyperion Castelnau, 1834
   * Megamorio Chaudoir, 1880
   * Morion Latreille, 1810
   * Morionidius Chaudoir, 1880
   * Moriosomus Motschulsky, 1864
   * Platynodes Westwood, 1846
   * Stereostoma Murray, 1857
* Tribù Pterostichini Bonelli, 1810
   * Sottotribù Euchroina Chaudoir, 1874
      * Abacillius Straneo, 1949
      * Abacillodes Straneo, 1988
      * Abaris Dejean, 1831
      * Apsaustodon Tschitscherine, 1901
      * Argutoridius Chaudoir, 1876
      * Blennidus Motschulsky, 1865
      * Blenniventer Straneo, 1986
      * Bothynoproctus Tschitscherine, 1900
      * Cephalostichus Straneo, 1977
      * Cynthidia Chaudoir, 1873
      * Euchroa Brulle, 1834
      * Eumara Tschitscherine, 1901
      * Gastrogmus Sloane, 1915
      * Haplobothynus Tschitscherine, 1901
      * Hybothecus Chaudoir, 1874
      * Litarthrum Sloane, 1915
      * Lobobrachus Sharp, 1885
      * Marsyas Putzeys, 1845
      * Meropalpus Tschitscherine, 1900
      * Microcephalus Dejean, 1828
      * Mireius Straneo, 1984
      * Nelidus Chaudoir, 1878
      * Neotalus Will, 2002
      * Oribazus Chaudoir, 1874
      * Pachythecus Chaudoir, 1874
      * Phaenaulax Tschitscherine, 1898
      * Prosopogmus Chaudoir, 1865
      * Pseudabarys Chaudoir, 1873
      * Setalimorphus Sloane, 1895
      * Setalis Castelnau, 1867
      * Simodontus Chaudoir, 1843
      * Trirammatus Chaudoir, 1838
   * Sottotribù Loxandrina Erwin & Sims, 1984
      * Adrimus Bates, 1871
      * Cerabilia Castelnau, 1867
      * Feronista Moore, 1965
      * Haploferonia Darlington, 1962
      * Homalonesiota Maindron, 1908
      * Loxandrus LeConte, 1852
      * Metoncidus Bates, 1871
      * Nebrioferonia Straneo, 1939
      * Oxycrepis Reiche, 1843
      * Stolonis Motschulsky, 1866
      * Zeodera Castelnau, 1867
   * Sottotribù Metiina Straneo, 1951
      * Abropus Waterhouse, 1842
      * Antarctiola Straneo, 1977
      * Chaetauchenium Tschitscherine, 1900
      * Feroniola Tschitscherine, 1900
      * Kuschelinus Straneo, 1963
      * Metius Curtis, 1839
      * Parhypates Motschulsky, 1866
   * Sottotribù Molopina Bonelli, 1810
      * Abacops Tschitscherine, 1902
      * Abax Bonelli, 1810
      * Aristochroa Tschitscherine, 1898
      * Aristochroodes Marcillac, 1993
      * Camptoscelis Dejean, 1828
      * Cedrorum Borges & Serrano, 1993
      * Chalcochrous Chaudoir, 1838
      * Eucamptognathus Chaudoir, 1837
      * Eudromus Klug, 1835
      * Henrotius Jeannel, 1953
      * Lesticus Dejean, 1828
      * Molopidius Jeannel, 1942
      * Molopinus Jeannel, 1948
      * Molops Bonelli, 1810
      * Myas Sturm, 1826: 171
      * Nirmala Andrewes, 1930
      * Orthomus Chaudoir, 1838
      * Oscadytes Lagar Mascaro, 1975
      * Pedius Motschulsky, 1850
      * Percus Bonelli, 1810
      * Peyrieraselus Deuve, 1981
      * Sthenocranion Tschitscherine, 1899
      * Styracoderus Chaudoir, 1874
      * Teratotarsa Tschitscherine, 1893
      * Trigonognatha Motschulsky, 1858
      * Tropidocerus Chaudoir, 1878
      * Zariquieya Jeannel, 1924
   * Sottotribù Pterostichina sensu stricto
      * Abacoleptus Fauvel, 1903
      * Abacomorphus Chaudoir, 1878
      * Abacophrastus Will, 2011
      * Acanthoferonia Moore, 1965
      * Allotriopus Bates, 1882
      * Analoma Darlington, 1971
      * Aulacopodus Britton, 1940
      * Basilewskya Straneo, 1948
      * Castelnaudia Tschitscherine, 1890
      * Catadromus W.S. Macleay, 1825
      * Conchitella Moore, 1962
      * Cophosomorpha Tschitscherine, 1890
      * Cratoferonia Tschitscherine, 1902
      * Cratogaster Blanchard, 1843
      * Cyclotrachelus Chaudoir, 1838
      * Cyrtoderus Hope, 1842
      * Darodilia Castelnau, 1867
      * Delinius Westwood, 1864
      * Euryabax Fauvel, 1903
      * Euryaptus Bates, 1892
      * Gastrellarius Casey, 1918
      * Gourlayia Britton, 1964
      * Harpostomus Chaudoir, 1856
      * Holcaspis Chaudoir, 1865
      * Leiradira Castelnau, 1867
      * Liopasa Tschitscherine, 1901
      * Lophoglossus LeConte, 1852
      * Loxodactylus Chaudoir, 1865
      * Loxogenius Sloane, 1907
      * Mecynognathus W.J.MacLeay, 1873
      * Megadromus Motschulsky, 1865
      * Neoferonia Britton, 1940
      * Nesites Andrewes, 1939
      * Notabax Moore, 1976
      * Notolestus Sloane, 1894
      * Notonomus Chaudoir, 1862
      * Nurus Motschulsky, 1866
      * Oberthueria Vuillet, 1911
      * Ogmophora Tschitscherine, 1896
      * Onawea Johns, 2007
      * Pachymorphus Chaudoir, 1838
      * Paniestichus Will, 2011
      * Paranurus Tschitscherine, 1901
      * Pareuryaptus Dubault, Lassalle & Roux, 2008
      * Percolaus Bates, 1882
      * Piesmus LeConte, 1852
      * Platycaelus Blanchard, 1843
      * Platysmodes Fauvel, 1903
      * Plocamostethus Britton, 1940
      * Poecilinus Jeannel, 1948
      * Poecilus Bonelli, 1810
      * Psegmatopterus Chaudoir, 1878
      * Pseudoceneus Tschitscherine, 1891
      * Pterostichus Bonelli, 1810
      * Rhabdotus Chaudoir, 1865
      * Rhytiferonia Darlington, 1962
      * Rhytisternus Chaudoir, 1865
      * Sarticus Motschulsky, 1865
      * Secatophus Castelnau, 1867
      * Setalidius Chaudoir, 1878
      * Speluncarius Reitter, 1886
      * Speomolops Patrizi, 1955
      * Sphodrosomus Perroud & Montrouzier, 1864
      * Stereocerus Kirby, 1837
      * Steropanus Fairmaire, 1888
      * Stomis Clairville, 1806
      * Straneostichus Sciaky, 1994
      * Taphoxomimus Straneo, 1975
      * Tapinopterus Schaum, 1858
      * Teropha Castelnau, 1867
      * Trichosternus Chaudoir, 1865
      * Trigonotoma Dejean, 1828
      * Wahlbergiana Bousquet, 2002
      * Xenion Tschitscherine, 1902
      * Zeopoecilus Sharp, 1886
* Tribù Zabrini Bonelli, 1810
   * Sottotribù Amarina C. Zimmermann, 1831
      * Amara Bonelli, 1810
      * Pseudamara Lindroth, 1968
   * Sottotribù Zabrina Bonelli, 1810
      * Zabrus Clairville, 1806
