Harpalus
# Harpalus

Harpalus es un género de coleópteros adéfagos de la familia Carabidae.

Principalmente holártico con unas pocas especies en la región afrotropical. Hay 415 especies.

## Algunas especies

Comprende las siguientes especies:

* Harpalus actiosus Casey, 1914
* Harpalus acupalpoides Reitter, 1900
* Harpalus adenticulatus Huang, 1992
* Harpalus aeneipennis Faldermann, 1836
* Harpalus aenigma (Tschitscherine, 1897)
* Harpalus aequicollis Motschulsky, 1844
* Harpalus aesculanus Pantel, 1888
* Harpalus affinis (Schrank, 1781)
* Harpalus agakhaniantzi Mikhailov, 1972
* Harpalus akinini Tschitscherine, 1895
* Harpalus alajensis Tschitscherine, 1898
* Harpalus albanicus Reitter, 1900
* Harpalus alexandrae Kataev, 1990
* Harpalus alexeevi Kataev, 1990
* Harpalus alienus Bates, 1878
* Harpalus alpivagus Tschitscherine, 1899
* Harpalus amarellus Bates, 1891
* Harpalus amariformis Motschulsky, 1844
* Harpalus amplicollis Menetries, 1848
* Harpalus amputatus Say, 1830
* Harpalus anatolicus Tschitscherine, 1898
* Harpalus angulatus Putzeys, 1878
* Harpalus angustipennis Boheman, 1848
* Harpalus angustitarsis Reitter, 1887
* Harpalus animosus Casey, 1924
* Harpalus anisodactyliformis Solsky, 1874
* Harpalus antonowi Tschitscherine, 1898
* Harpalus anxioides Kataev, 1991
* Harpalus anxius (Duftschmid, 1812)
* Harpalus aogashimensis (Habu, 1957)
* Harpalus apache Kataev, 2010
* Harpalus araraticus Mlynar, 1979
* Harpalus arcuatus Tschitscherine, 1898
* Harpalus arnoldii Kataev, 1988
* Harpalus ascetes Kataev & Wrase, 1997
* Harpalus asemus Basilewsky, 1946
* Harpalus asphaltinus Roth, 1851
* Harpalus aterrimus Casey, 1914
* Harpalus atratus Latreille, 1804
* Harpalus atrichatus Hatch, 1949
* Harpalus atripes Casey, 1914
* Harpalus attenuatus Stephens, 1828
* Harpalus australasiae Dejean, 1829
* Harpalus autumnalis Duftschmid, 1812
* Harpalus azumai Habu, 1968
* Harpalus babai Habu, 1973
* Harpalus badakschanus Jedlicka, 1956
* Harpalus baleensis Clarke, 1973
* Harpalus balli Noonan, 1991
* Harpalus basanicus J.R. Sahlberg, 1911
* Harpalus basilewskyi Facchini, 2003
* Harpalus basuto Basilewsky, 1958
* Harpalus bellieri Reiche, 1861
* Harpalus beneshi Kataev & Wrase, 1997
* Harpalus bitinctus Jeannel, 1948
* Harpalus brachypterus Tschitcherine, 1898
* Harpalus brevicornis Germar, 1824
* Harpalus brevis Motschiulsky, 1844
* Harpalus breviusculus Chaudoir, 1846
* Harpalus brunnipes Dejean, 1829
* Harpalus bucharicus Tschitscherine, 1898
* Harpalus bungii Chaudoir, 1844
* Harpalus caeruleatus Bates, 1878
* Harpalus caiphus Reiche & Saulcy, 1855
* Harpalus calathoides Motschulsky, 1844
* Harpalus calceatus (Duftschmid, 1812)
* Harpalus caliginosus (Fabricius, 1775)
* Harpalus capicola Dejean, 1829
* Harpalus capito A.Morawitz, 1862
* Harpalus cardoni Antoine, 1922
* Harpalus caspius (Steven, 1806)
* Harpalus cautus Dejean, 1829
* Harpalus cephalotes Fairmaire & Laboulbene, 1854
* Harpalus chalcentus Bates, 1873
* Harpalus chalcopterus Jeannel, 1948
* Harpalus chasanensis Lafer, 1989
* Harpalus chengjiangensis Huang, 1993
* Harpalus chiloschizontus Huang, 1995
* Harpalus chobautianus Lutshnik, 1922
* Harpalus chrysopus Reitter, 1887
* Harpalus cilihumerus Huang, Hu & Sun, 1994
* Harpalus circumpunctatus Chaudoir, 1846
* Harpalus cisteloides Motschulsky, 1844
* Harpalus cohni Ball, 1972
* Harpalus colchicus Motschulsky, 1850
* Harpalus compar Leconte, 1848
* Harpalus compressus Motschulsky, 1844
* Harpalus contemptus Dejean, 1829
* Harpalus convexus Faldermann, 1836
* Harpalus cordifer Notman, 1919
* Harpalus cordifer Notman, 1919
* Harpalus coreanus (Tschitscherine, 1895)
* Harpalus corporosus Motschulsky, 1861
* Harpalus corpulentus (Casey, 1914)
* Harpalus corrugatus Basilewsky, 1958
* Harpalus crates Bates, 1883
* Harpalus cupreus Dejean, 1829
* Harpalus cursorius Peringuey, 1896
* Harpalus cyanopterus Tschitscherine, 1897
* Harpalus cyclogonus Chaudoir, 1844
* Harpalus cyrtonotoides Notman, 1919
* Harpalus danieli Reitter, 1900
* Harpalus davidi (Tschitscherine, 1897)
* Harpalus davidianus Tschitscherine, 1903
* Harpalus decipiens Dejean, 1829
* Harpalus defector Peringuey, 1896
* Harpalus demelti Korge, 1962
* Harpalus desertus Leconte, 1859
* Harpalus diligens Tschitscherine, 1898
* Harpalus dimidiatus (P. Rossi, 1790)
* Harpalus disaogashimensis Huang In Huang & Zhang, 1995
* Harpalus disimuciulus Huang, Lei, Yan & Hu, 1996
* Harpalus dispar Dejean, 1829
* Harpalus dissitus Antoine, 1931
* Harpalus distinguendus (Duftschmid, 1812)
* Harpalus diversicollis Basilewsky, 1958
* Harpalus dubius Boheman, 1848
* Harpalus durangoensis Bates, 1891
* Harpalus ebeninus Heyden, 1870
* Harpalus eberlovi O. Berlov, 1996
* Harpalus egenus Dejean, 1829
* Harpalus egorovi Lafer, 1989
* Harpalus ellipsis Leconte, 1848
* Harpalus eous Tschitscherine, 1901
* Harpalus eraticus Say, 1823
* Harpalus erosus Mannerheim, 1825
* Harpalus erythropus Dejean, 1829
* Harpalus euchlorus Menetries, 1839
* Harpalus exiguus Boheman, 1848
* Harpalus famelicus Tschitscherine, 1898
* Harpalus farkaci Kataev & Wrase, 1995
* Harpalus faunus Say, 1823
* Harpalus ferghanensis Tschitscherine, 1899
* Harpalus fimetarius Dejean, 1829
* Harpalus flavescens Piller & Mitterpacher, 1783
* Harpalus flavicornis Dejean, 1829
* Harpalus fokienensis Schauberger, 1930
* Harpalus foveiger Tschitscherine, 1895
* Harpalus franzi Mateu, 1954
* Harpalus frater Chaudoir, 1876
* Harpalus fraternus Leconte, 1852
* Harpalus froelichii Sturm, 1818
* Harpalus fukiensis Jedlicka, 1956
* Harpalus fulgens Csiki, 1932
* Harpalus fulvicornis Thunberg, 1806
* Harpalus fulvilabris Mannerheim, 1853
* Harpalus fulvipennis Chaudoir, 1843
* Harpalus fulvus Dejean, 1829
* Harpalus fuscicornis Menetries, 1832
* Harpalus fuscipalpis Sturm, 1818
* Harpalus fuscipennis Wiedemann, 1825
* Harpalus fuscoaeneus Dejean, 1829
* Harpalus ganssuensis Semenov, 1889
* Harpalus giacomazzoi Kataev & Wrase, 1996
* Harpalus gilgil Basilewsky, 1946
* Harpalus gisellae Csiki, 1932
* Harpalus glasunovi Kataev, 1987
* Harpalus gravis Leconte, 1858
* Harpalus gregoryi Alluaud, 1917
* Harpalus grilli Kataev, 2001
* Harpalus griseus (Panzer, 1796)
* Harpalus hartmanni Kataev, 2001
* Harpalus hatchi Ball & Anderson, 1962
* Harpalus hauserianus Schauberger, 1929
* Harpalus herbivagus Say, 1823
* Harpalus heyrovskyi Jedlicka, 1928
* Harpalus hiekei Kataev & Wrase, 2010
* Harpalus hirtipes Panzer, 1796
* Harpalus honestus (Dufischmid, 1812)
* Harpalus hospes Sturm, 1818
* Harpalus huachuca Ball, 1972
* Harpalus hybridus Boheman, 1848
* Harpalus hypogeomysis Huang, 1993
* Harpalus idiotus Bates, 1889
* Harpalus imerinae Jeannel, 1948
* Harpalus impressicollis Jeannel, 1948
* Harpalus impressus Roth, 1851
* Harpalus inconcinnus Chaudoir, 1876
* Harpalus indianus Csiki, 1932
* Harpalus indicola Bates, 1878
* Harpalus indicus Bates, 1891
* Harpalus indigens Casey, 1924
* Harpalus inexspectatus Kataev, 1989
* Harpalus ingenuus Tschitscherine, 1898
* Harpalus innocuus Leconte, 1863
* Harpalus italus Schaum, 1860
* Harpalus janinae Jeanne, 1984
* Harpalus jeanneli Basilewsky, 1946
* Harpalus jordanus Jedlicka, 1964
* Harpalus jureceki (Jedlicka, 1928)
* Harpalus kabakianus Kataev, 1988
* Harpalus kabakovi Kataev, 1987
* Harpalus kadleci Kataev & Wrase, 1995
* Harpalus kadyrbekovi Kataev, 1988
* Harpalus kagyzmanicus Kataev, 1984
* Harpalus kailiensis Huang, 1992
* Harpalus kandaharensis Jedlicka, 1956
* Harpalus karamani Apfelbeck, 1902
* Harpalus karokorum Jedlicka, 1958
* Harpalus katiae F.Battoni, 1985
* Harpalus kazanensis Jedlicka, 1958
* Harpalus kaznakovi Kataev & Wrase, 1997
* Harpalus kibonoti Alluaud, 1926
* Harpalus kirgisicus Motschulsky, 1844
* Harpalus kiritshenkoi Kataev, 1990
* Harpalus klapperichi Jedlicka, 1956
* Harpalus kmecoi Facchini, 2003
* Harpalus koshiensis Kirschenhofer, 1992
* Harpalus kozlovi Kataev, 1993
* Harpalus kryzhanovskii Kataev, 1988
* Harpalus kunarensis Kataev, 1993
* Harpalus laetus Reiche, 1843
* Harpalus laevipes Zetterstedt, 1828
* Harpalus lama Kataev & Wrase, 1997
* Harpalus lampronotus Jeannel, 1948
* Harpalus lateralis Dejean, 1829
* Harpalus laticeps Leconte, 1850
* Harpalus latus (Linnaeus, 1758)
* Harpalus lederi Tschitscherne, 1899
* Harpalus lethierryi Reiche, 1860
* Harpalus lewisii Leconte, 1865
* Harpalus liobasis Chaudoir, 1868
* Harpalus litigiosus Dejean, 1829
* Harpalus longicollis Leconte, 1848
* Harpalus longihornus Lei & Huang, 1997
* Harpalus longipalmatus Mordkovitsh, 1969
* Harpalus lopezi Serrano & Lencina, 2008
* Harpalus lugubris Boheman, 1848
* Harpalus lumbaris Mannerheim, 1825
* Harpalus luteicornis Dufischmid, 1812
* Harpalus lutshniki Schauberger, 1932
* Harpalus macronotus Tschitscherine, 1893
* Harpalus madagascariensis Dejean, 1831
* Harpalus mairei Peyerimhoff, 1928
* Harpalus major Motschulsky, 1850
* Harpalus makhekensis Basilewsky, 1958
* Harpalus manas Kataev, 1990
* Harpalus marginellus Gyllenhal, 1827
* Harpalus mariae Kataev, 1997
* Harpalus martini Vandyke, 1926
* Harpalus masoreoides Bates, 1878
* Harpalus massarti Burgeon, 1935
* Harpalus mauritanicus Gaubil, 1844
* Harpalus medvedevi Kataev, 2006
* Harpalus megacephalus Leconte, 1848
* Harpalus meghalayensis Kataev, 2001
* Harpalus melancholicus Dejean, 1829
* Harpalus melaneus Bates, 1878
* Harpalus meridianus Andrewes, 1923
* Harpalus metallinus Menetries, 1836
* Harpalus metarsius Andrewes, 1930
* Harpalus meteorus Basilewsky, 1946
* Harpalus michaili Kataev, 1990
* Harpalus michailovi Kataev, 1987
* Harpalus microdemas Schauberger, 1932
* Harpalus microthorax Motschulsky, 1849
* Harpalus miles Peringuey, 1896
* Harpalus minutulus Kataev & Liang, 2004
* Harpalus mitridati Pliginskiy, 1915
* Harpalus mlynari Kataev, 1990
* Harpalus modestus Dejean, 1829
* Harpalus morvani Kataev, 2001
* Harpalus muciulus Huang, 1992
* Harpalus nanniscus Peringuey, 1896
* Harpalus natalensis Boheman, 1848
* Harpalus natalicus Peringuey, 1896
* Harpalus neglectus Audinet-Serville, 1821
* Harpalus nevadensis K. Daniel & J. Daniel, 1898
* Harpalus nigrans A. Morawitz, 1862
* Harpalus nigritarsis C.R. Sahlberg, 1827
* Harpalus numidicus Bedel, 1893
* Harpalus nyassicus Basilewsky, 1946
* Harpalus obliquus G.Horn, 1880
* Harpalus oblitus Dejean, 1829
* Harpalus obnixus Casey, 1924
* Harpalus obtusiusculus Jeannel, 1948
* Harpalus ochropus Kirby, 1837
* Harpalus oodioides Dejean, 1829
* Harpalus opacipennis (Haldemann, 1843)
* Harpalus optabilis Dejean, 1829
* Harpalus ovtshinnikovi Kataev, 1990
* Harpalus pachys Jeannel, 1948
* Harpalus paivanus Wollaston, 1867
* Harpalus pallidipennis A. Morawitz, 1862
* Harpalus pallipes Chaudoir, 1837
* Harpalus parasinuatus Kataev & Liang, 2007
* Harpalus paratus Casey, 1924
* Harpalus parvulus Dejean, 1829
* Harpalus pastor Motschulsky, 1844
* Harpalus penglainus Huang, Hu & Sun, 1994
* Harpalus pensylvanicus (Degeer, 1774)
* Harpalus periglabellus Huang, 1992
* Harpalus perlucens Bates, 1878
* Harpalus petri Tschitscherine, 1902
* Harpalus picipennis Duftschmid, 1812
* Harpalus pilosus Huang, 1995
* Harpalus plancyi Tschitscherine, 1897
* Harpalus platynotus Bates, 1873
* Harpalus plenalis Casey, 1914
* Harpalus politus Dejean, 1829
* Harpalus polyglyptus Schaum, 1862
* Harpalus poncei Will, 2001 Usa (Florida)
* Harpalus potanini Tschitscherine, 1906
* Harpalus poussereaui Pierre & Mauro, 2010
* Harpalus praecurrens Schauberger, 1934
* Harpalus praticola Bates, 1891
* Harpalus procognatus Lorenz, 1998
* Harpalus progrediens Schauberger, 1922
* Harpalus prosperus Basilewsky, 1972
* Harpalus protractus Casey, 1914
* Harpalus providens Casey, 1914
* Harpalus pseudohauserianus Kataev, 2001
* Harpalus pseudophonoides Schauberger, 1930
* Harpalus pseudoserripes Reitter, 1900
* Harpalus pseudotinctulus Schauberger, 1932
* Harpalus pterostichus Reitter, 1900
* Harpalus puetzi Kataev & Wrase, 1997
* Harpalus pulchirinulus Reitter, 1900
* Harpalus pulvinatus Menetries, 1848
* Harpalus pumilus Sturm, 1818
* Harpalus punctatostriatus Dejean, 1829
* Harpalus puncticeps (Casey, 1914)
* Harpalus punctipennis Mulsant, 1852
* Harpalus pusillus Motschulsky, 1850
* Harpalus pygmaeus Dejean, 1829
* Harpalus quadratus Chaudoir, 1846
* Harpalus raphaili Kataev, 1997
* Harpalus reflexus Putzeys, 1878
* Harpalus regressus (Casey, 1914)
* Harpalus remboides Solsky, 1874
* Harpalus retractus Leconte, 1863
* Harpalus reversus Casey, 1924
* Harpalus rivalsi Jeannel, 1948
* Harpalus roninus Bates, 1873
* Harpalus rotundus Facchini, 2003
* Harpalus rougemonti Clarke, 1973
* Harpalus rubefactus Bates, 1873
* Harpalus rubripes Dufischmid, 1812
* Harpalus rufipalpis Sturm, 1818
* Harpalus rufipes (Degeer, 1774)
* Harpalus rufiscapus Gebler, 1833
* Harpalus rufitarsoides Schauberger, 1934
* Harpalus rufomarginatus (Boheman, 1848)
* Harpalus rumelicus Apfelbeck, 1904
* Harpalus salinulus Reitter, 1900
* Harpalus salinus Dejean, 1829
* Harpalus sanctaehelenae Basilewsky, 1972
* Harpalus saxicola Dejean, 1829
* Harpalus scabripectus Huang, Hu & Sun, 1994
* Harpalus schaumii Wollaston, 1864
* Harpalus semenowi Tschitscherine, 1901
* Harpalus sericatus Tschitscherine, 1906
* Harpalus serripes Quensel, 1806
* Harpalus servus Duftschmid, 1812
* Harpalus setiporus Reitter, 1894
* Harpalus seyrigi Jeannel, 1948
* Harpalus siculus Dejean, 1829
* Harpalus signaticornis (Duftschmid, 1812)
* Harpalus silipes Dejean, 1831
* Harpalus simplicidens Schauberger, 1929
* Harpalus singularis Tschitscherine, 1906
* Harpalus sinicus Hope, 1845
* Harpalus sinuatipennis Jeannel, 1948
* Harpalus sinuatus Tschitscherine, 1893
* Harpalus smaragdinus Duftschmid, 1812
* Harpalus smyrnensis Heyden, 1888
* Harpalus solitaris Dejean, 1829
* Harpalus somereni Basilewsky, 1946
* Harpalus somnulentus Dejean, 1829
* Harpalus spadiceus Dejean, 1829
* Harpalus spretus Peringuey, 1896
* Harpalus spurius Peringuey, 1896
* Harpalus stephani Ball, 1972
* Harpalus stevenii Dejean, 1829
* Harpalus stoetznerianus Schauberger, 1932
* Harpalus strenuus Tschitscherine, 1898
* Harpalus stricticollis Jeannel, 1948
* Harpalus subaeneus Boheman, 1848
* Harpalus subcylindricus Dejean, 1829
* Harpalus subphaedrus Basilewsky, 1946
* Harpalus subtruncatus Chaudoir, 1846
* Harpalus suensoni Kataev, 1997
* Harpalus sulphuripes Germar, 1824
* Harpalus sundaicus Schauberger, 1933
* Harpalus sushenicus Kataev, 1990
* Harpalus suturangulus Reitter, 1887
* Harpalus szalliesi Kataev & Wrase, 1995
* Harpalus taciturnus Dejean, 1829
* Harpalus tadorcus Ball, 1972
* Harpalus tangutorum Kataev, 1993
* Harpalus tardus Panzer, 1796
* Harpalus tarsalis Mannenheim, 1825
* Harpalus tenebrosus Dejean, 1829
* Harpalus terrestris Motschulsky, 1844
* Harpalus texanus Casey, 1914
* Harpalus tianschanicus Semenov, 1889
* Harpalus tibeticus Andrewes, 1930
* Harpalus tichonis Jakobson, 1907
* Harpalus tinctulus Bates, 1873
* Harpalus tiridates Reitter, 1900
* Harpalus tithonus Reitter, 1900
* Harpalus torosensis Jedlicka, 1961
* Harpalus torridoides Reitter, 1900
* Harpalus trichophorus Tschitscherine, 1897
* Harpalus tridens A.Morawitz, 1862
* Harpalus triseriatus A. Fleischer, 1897
* Harpalus turcicus Jedlicka, 1958
* Harpalus turmalinus Erichson, 1847
* Harpalus udege Lafer, 1989
* Harpalus uniformis Motschulsky, 1844
* Harpalus ussuricus Mlynar, 1979
* Harpalus ussuriensis Chaudoir, 1863
* Harpalus vanemdeni Schauberger, 1932
* Harpalus venator Boheman, 1848
* Harpalus ventralis Leconte, 1848
* Harpalus vereschaginae Kataev, 1988
* Harpalus vernicosus Kataev & Liang, 2007
* Harpalus viridanus Motschulsky, 1844
* Harpalus viridicupreus Reiche, 1843
* Harpalus vittatus Gebler, 1833
* Harpalus wadiensis Jedlicka, 1964
* Harpalus wohlberedti Emden & Schauberger, 1932
* Harpalus xanthopus Gemminger & Harold, 1868
* Harpalus xinjiangensis Huang, Hu & Sun, 1994
* Harpalus yinchuanensis Huang, 1993
* Harpalus zabroides Dejean, 1829
* Harpalus zhdankoi Kataev, 1990
