Assistant (1956), Head of Works (1958) and Assistant Professor (1962) at the Faculty of Sciences of Paris, Doctor of Science (1963) after his return from military service, Professor in Tunis [scientific and technical cooperation] (1963-1966) then in Rouen (1966-1994) where he directed the Laboratory associated with the CNRS Cell Physiology, Signals and Regulations (1978-1990).