Of that number, Žerjavić estimated that 78,000 were killed by the Ustaše at Jasenovac and in "prisons, pits and other camps", 45,000 were killed by German forces, 15,000 by Italian forces, 34,000 were killed in battles between the Ustaše, the Chetniks, and the Partisans, and 25,000 died of typhoid.