With 80% fossils attributed to him, le Rhinolophidae, Protorhinolophus shanghuangensis, dominates the community of bats Shanghuang. Its dental morphology combines characteristics relating to Rhinolophidae (for example, the presence of a third lower premolar vestigial) with characters that are found in the most primitive forms, underscoring its archaic appearance. It is also possible to find morphological similarities with another kind fossil discovered in karst fillings Quercy (South-West of France) dating from the late Middle Eocene (about -38 million years).
Na 80% fossils kuhusishwa kwake, le Rhinolophidae, Protorhinolophus shanghuangensis, dominates jamii ya popo Shanghuang . Morphology wake meno unachanganya sifa zinazohusiana na Rhinolophidae (kwa mfano, uwepo wa tatu chini premolar vestigial) na wahusika kwamba ni kupatikana katika aina nyingi za primitive, akisisitiza muonekano wake kizamani . Ni pia inawezekana kupata kufanana kubadilika kwa maumbile na mafuta mwingine aina aligundua katika karst fillings Quercy (South-West ya Ufaransa) yaliyoanza mwishoni mwa Mashariki ya Eocene (kuhusu -38 miaka milioni).